Tortula protobryoides

R. H. Zander

Bull. Buffalo Soc. Nat. Sci. 32: 226. 1993,.

Basionyms: Phascum bryoides Dickson Fasc. Pl. Crypt. Brit. 4: 3, plate 10, fig. 3. 1801,
Synonyms: Pottia bryoides (Dickson) Mitten Protobryum bryoides (Dickson) J. Guerra & M. J. Cano
Treatment appears in FNA Volume 27. Treatment on page 592. Mentioned on page 587, 589, 629.

Leaves ovate to oblong-lanceolate, apex broadly acute to rounded, mucronate to short-awned, margins revolute in mid 2/3–3/4 of leaf, weakly bordered from apex to near base with (2–) 4 rows of weakly thicker-walled and less-papillose or smooth cells; costa excurrent, lacking an adaxial pad of cells though convex adaxially, distally narrow, 3–4 cells across adaxial surface; distal laminal cells subquadrate to hexagonal, occasionally somewhat elongated longitudinally, width 13–18 µm wide, 1 (–2):1, papillose with 4 (–6) simple papillae per lumen. Sexual condition autoicous. Sporophytes exerted. Seta length 0.2–0.4 cm. Capsule stegocarpic, not systylius, broadly cylindric, erect and nearly straight, urn ca. 1.4–1.5 mm; peristome remaining adherent to the inside of the operculum, teeth rudimentary to linear, to 300 µm, of several filaments variously anastomosing and weakly twisted, basal membrane length to 50 µm; operculum ca. 0.7–0.8 mm, not falling. Spores ca. 20–25 µm, spheric, densely low-papillose.


Phenology: Capsules mature winter–spring.
Habitat: Soil
Elevation: low elevations

Distribution

V27 846-distribution-map.gif

B.C., Sask., Ariz., Calif., Colo., Minn., Mexico (Baja California), Europe, sw Asia

Discussion

Selected References

None.

Lower Taxa

None.

"um" is not declared as a valid unit of measurement for this property.

... more about "Tortula protobryoides"
smooth +  and papillose +
smooth +  and papillose +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
channeled +, nearly flat +  and concave +
0.1 cm1 mm <br />0.001 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
of 1-2 rows +  and differentiated +
long-excurrent +  and short-excurrent +
Richard H. Zander +  and Patricia M. Eckel +
R. H. Zander +
semicircular +  and elliptical +
semicircular +, elliptical +  and rounded +
papillose +  and smooth +
usually slightly wider +
usually ovate;oblong +
differentiated +
Phascum bryoides +
obovoid +
cucullate +
0.25 cm2.5 mm <br />0.0025 m <br /> (?) +  and 0.6 cm6 mm <br />0.006 m <br /> (?) +
not systylius +
0.3 cm3 mm <br />0.003 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
dark-brown +  and yellowish-brown +
cleistocarpous +  and stegocarpous +
cylindric +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
monoicous +, dioicous +  and sexual +
thicker-walled +
1 (?) +  and 3 (?) +
smooth +  and less-papillose +
elongate-rhomboid +  and elongate +
tomentose +  and bare +
subpercurrent +
autoicous +  and sexual +
B.C. +, Sask. +, Ariz. +, Calif. +, Colo. +, Minn. +, Mexico (Baja California) +, Europe +  and sw Asia +
low elevations +
differentiated +
multicellular +
somewhat larger +
yellow +  and orange-red or red +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
concave;keeled;channeled;ovoid;lanceolate or lingulate +
ovate +  and oblong-lanceolate +
reduced +
serrulate +  and entire +
smaller +
rostrate;long-conic +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
differentiated +
photosynthetic +
lateral +  and terminal +
lateral +  and terminal +
red +, orange +  and yellow +
consisting of 16 mostly 2 times cleft spiculose striate or papillose lanceolate teeth , , , , +  and rudimentary +
Capsules mature winter–spring. +
Bull. Buffalo Soc. Nat. Sci. +
straight +
not twisted +  and twisted +
brown;yellowish-brown +
0.2 cm2 mm <br />0.002 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
elongate +
0 cm0 mm <br />0 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
2 +  and 1 +
orange +, red +, yellow +, laminal +  and light-brown +
spheric +
rounded-pentagonal +
0 cm0 mm <br />0 m <br /> (2 cm20 mm <br />0.02 m <br />) +
Pottia bryoides +  and Protobryum bryoides +
Tortula protobryoides +
species +
ovoid;cylindric +
rudimentary +
32 +  and 16 +
triangular +
0.14 cm1.4 mm <br />0.0014 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
yellow distally and red +, orange +, red +  and yellow +
rectangular +
thickened +, thin +  and thicker +
stemless +  and perigoniate +
yellowbrown;dark-brown +
occasionally smaller than the perichaetiate +