Tortula systylia

(Schimper) Lindberg

Musc. Scand., 20. 1879,.

Basionyms: Desmatodon systylius Schimper Flora 28: 145. 1845
Treatment appears in FNA Volume 27. Treatment on page 599. Mentioned on page 588, 590, 600.

Leaves obovate to occasionally ovate, apex acute or broadly acute, awned, margins recurved or plane proximally, not bordered; costa excurrent into the awn, lacking an adaxial pad of cells, distally narrow, 2–3 cells across the convex adaxial surface; distal laminal cells hexagonal to rhomboidal, width 15–22 µm wide, 1: 1 near apex but elongate at mid leaf and basally, smooth. Sexual condition autoicous. Sporophytes exerted. Seta 0.8–1.1 cm. Capsule systylius, short-cylindric, erect and nearly straight, urn 1.5–2 mm; peristome 200–250 µm, teeth 16, straight, divided to near base but variously fused or perforate, basal membrane low; operculum 0.5–0.7 mm. Spores 22–30 (–35) µm, spheric, warty papillose.


Phenology: Capsules mature summer.
Habitat: Soil in rock crevices, tundra
Elevation: high elevations (0–3700 m)

Distribution

V27 860-distribution-map.gif

Greenland, Alta., B.C., Man., Nfld. and Labr. (Nfld.), N.W.T., Nunavut, Que., Yukon, Alaska, Calif., Colo., Wyo., Europe, Asia

Discussion

Tortula systylia may be confused with Stegonia but the latter has a distinct triangle of echlorophyllose cells at the apex of the much broader leaves. It is similar also to T. hoppeana, but the laminal cells are smooth. After dehiscence, the systylius capsules retain a raised operculum on the columellar stalk.

Selected References

None.

Lower Taxa

None.

"um" is not declared as a valid unit of measurement for this property."um" is not declared as a valid unit of measurement for this property.

... more about "Tortula systylia"
smooth +  and papillose +
smooth +  and papillose +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
of 1-2 rows +  and differentiated +
long-excurrent +  and short-excurrent +
Elongate (?) +  and Acute (?) +
Richard H. Zander +  and Patricia M. Eckel +
(Schimper) Lindberg +
semicircular +  and elliptical +
semicircular +, elliptical +  and rounded +
papillose +  and smooth +
usually slightly wider +
usually ovate;oblong +
differentiated +
Desmatodon systylius +
obovoid +
cucullate +
0.25 cm2.5 mm <br />0.0025 m <br /> (?) +  and 0.6 cm6 mm <br />0.006 m <br /> (?) +
0.3 cm3 mm <br />0.003 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
dark-brown +  and yellowish-brown +
cleistocarpous +  and stegocarpous +
short-cylindric +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
monoicous +, dioicous +  and sexual +
twisted +, vesiculose +  and thicker-walled +
1 (?) +  and 3 (?) +
elongate-rhomboid +  and elongate +
tomentose +  and bare +
subpercurrent +
autoicous +  and sexual +
Greenland +, Alta. +, B.C. +, Man. +, Nfld. and Labr. (Nfld.) +, N.W.T. +, Nunavut +, Que. +, Yukon +, Alaska +, Calif. +, Colo. +, Wyo. +, Europe +  and Asia +
high elevations (0–3700 m) +
differentiated +
multicellular +
Soil in rock crevices, tundra +
somewhat larger +
yellow +  and orange-red or red +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
concave;keeled;channeled;ovoid;lanceolate or lingulate +
obovate +  and occasionally ovate +
reduced +
serrulate +  and entire +
smaller +
rostrate;long-conic +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.07 cm0.7 mm <br />7.0e-4 m <br />) +
differentiated +
photosynthetic +
simple +  and hollow +
4 +  and 6 +
lateral +  and terminal +
lateral +  and terminal +
red +, orange +  and yellow +
consisting of 16 mostly 2 times cleft spiculose striate or papillose lanceolate teeth , , , , +  and rudimentary +
Capsules mature summer. +
Musc. Scand., +
straight +
not twisted +  and twisted +
brown;yellowish-brown +
elongate +
0.8 cm8 mm <br />0.008 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
2 +  and 1 +
orange +, red +, yellow +, laminal +  and light-brown +
spheric +
rounded-pentagonal +
0 cm0 mm <br />0 m <br /> (2 cm20 mm <br />0.02 m <br />) +
Desmatodon +, Phascum +, Pottia +  and Protobryum +
Tortula systylia +
species +
ovoid;cylindric +
straight +
rudimentary +
divided +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
yellow distally and red +, orange +, red +  and yellow +
rectangular +
thickened +, thin +  and thicker +
stemless +  and perigoniate +
yellowbrown;dark-brown +
occasionally smaller than the perichaetiate +