Transberingia bursifolia

(de Candolle) Al-Shehbaz & O’Kane

Novon 13: 396. 2003.

Basionym: Nasturtium bursifolium de Candolle Syst. Nat. 2: 194. 1821
Synonyms: Arabidopsis bursifolia (de Candolle) Botschantzev Beringia bursifolia (de Candolle) R. A. Price, Al-Shehbaz & O’Kane
Treatment appears in FNA Volume 7. Treatment on page 456.

Plants densely hirsute basally, trichomes primarily simple, to 1 mm, these often mixed with smaller, forked, and/or dendritic ones, glabrescent or sparsely pubescent distally. Stems simple or few to several from caudex, (0.5–) 1–4.5 (–6.5) dm. Basal leaves: petiole 0.3–3 cm, flattened at base, ciliate; blade oblanceolate to obovate, or linear-lanceolate, (0.5–) 1.2–6.5 (–10) cm × (2–) 3–8 (–17) mm, base cuneate to attenuate, apex obtuse to subacute, surfaces pubescent, trichomes usually branched. Cauline leaves: blade lanceolate to linear-lanceolate or oblong, (0.5–) 1–3 (–4.2) cm × (1–) 2.5–7 (–10) mm, base strongly sagittate. Fruiting pedicels not appressed to rachis, straight, (3–) 5–14 (–18) mm, glabrous. Flowers: sepals 1.2–2.5 × 0.6–1 mm, sparsely to densely pubescent; petals (2.5–) 3–4 (–4.5) × (0.8–) 1–1.5 mm; filaments 2–2.5 mm; anthers 0.4–0.5 mm. Fruits (1.5–) 2–3.3 (–4) cm × (0.7–) 1–1.5 (–2) mm; style slender, 0.2–0.8 (–1) mm. Seeds 0.8–1 × 0.4–0.5 mm.

Distribution

V7 696-distribution-map.gif

Alta., N.W.T., Nunavut, Sask., Yukon, Alaska, Calif., Colo., Idaho, Mont., Nev., Utah, Wyo., e Asia (Russian Far East)

Discussion

Subspecies 2 (2 in the flora).

R. C. Rollins (1993) treated subspp. bursifolia and virgata as distinct species of Halimolobos, but the alleged differences (stems branched basally versus distally, plants cespitose versus non-cespitose, fruits slightly compressed versus terete) do not hold, and the only reliable difference between the two taxa is in the type of trichomes. Therefore, a subspecific, rather than specific, rank is adopted here.

Selected References

None.

Key

1 Plants glabrous or sparsely pubescent on stems distally and on rachises, trichomes simple, spreading. Transberingia bursifolia subsp. bursifolia
1 Plants usually pubescent, rarely glabrescent, on stems distally and on rachises, trichomes branched, subappressed, these sometimes mixed with simple ones. Transberingia bursifolia subsp. virgata

"elongated" is not a number."thick" is not a number."dm" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property.

... more about "Transberingia bursifolia"
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
obtuse +  and subacute +
Ihsan A. Al-Shehbaz +
(de Candolle) Al-Shehbaz & O’Kane +
sagittate +, cuneate +  and attenuate +
Nasturtium bursifolium +
appendaged +  and unappendaged +
3 cm30 mm <br />0.03 m <br /> (4.2 cm42 mm <br />0.042 m <br />) +
0.7 cm7 mm <br />0.007 m <br /> (1 cm10 mm <br />0.01 m <br />) +
not +  and auriculate +
1 cm10 mm <br />0.01 m <br /> (3 cm30 mm <br />0.03 m <br />) +
lanceolate;linear-lanceolate or oblong +
much smaller +  and reduced +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
urceolate +, campanulate +  and tubular +
branched +  and simple +
subsessile +, petiolate +  and sessile +
well-developed +
distinct +
differentiated +
emarginate +  and entire +
incumbent +  and accumbent +
Alta. +, N.W.T. +, Nunavut +, Sask. +, Yukon +, Alaska +, Calif. +, Colo. +, Idaho +, Mont. +, Nev. +, Utah +, Wyo. +  and e Asia (Russian Far East) +
straight +  and curved +
not winged +  and unappendaged +
connate +  and distinct +
0.2 cm2 mm <br />0.002 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
actinomorphic +
2-valved +  and capsular +
3.3 cm33 mm <br />0.033 m <br /> (4 cm40 mm <br />0.04 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
2 cm20 mm <br />0.02 m <br /> (3.3 cm33 mm <br />0.033 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
latiseptate +  and unsegmented +
indehiscent +  and dehiscent +
angustiseptate +  and terete +
subsessile +  and sessile +
sessile +  and petiolate +
alternate +, not +  and rosulate +
cauline +  and basal +
connate +  and distinct +
decurrent +
dentate +  and entire +
forked +
smaller +
tenuinucellate +, crassinucellate +  and bitegmic +
campylotropous +  and anatropous +
0.4 cm4 mm <br />0.004 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
rudimentary +
oblanceolate +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
flattened +
0.3 cm3 mm <br />0.003 m <br /> (3 cm30 mm <br />0.03 m <br />) +
trinucleate +  and 3(-11)-colpate +
subdivaricate +
slender +
1.4 cm14 mm <br />0.014 m <br /> (1.8 cm18 mm <br />0.018 m <br />) +
mucilaginous +
not +  and mucilaginous +
white +, black +, brown +  and yellow +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
persistent +  and caducous +
distinct +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
reduced +
latiseptate +, schizocarpic +, samaroid +  and lomentaceous +
indehiscent +  and dehiscent +
angustiseptate +, flat +, angled +, terete +, torulose +, not +, segmented +  and nutletlike +
tetradynamous +
few to several +  and simple +
procumbent +  and decumbent +
ascending +  and erect +
2-lobed +, entire +, conical +  and capitate +
subsessile +, sessile +  and petiolate +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
persistent +
distinct +
slender +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
Arabidopsis bursifolia +  and Beringia bursifolia +
Transberingia bursifolia +
Transberingia +
species +
branched +  and simple +
pubescent +  and glabrescent +
0 cm0 mm <br />0 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
coiled +, 1-7-veined +  and veined +
pubescent +  and glabrous +
anastomosing +
pubescent +  and glabrous +
rhizomatous +, taprooted +, scapose +  and not +
perennial +, biennial +  and annual +
aquatics +  and terrestrial +