Veronica anagallis-aquatica

Linnaeus

Sp. Pl. 1: 12. 1753.

Common names: Water speedwell véronique mouron-d'eau
Introduced
Synonyms: Veronica anagallis-aquatica var. terrea Farwell V. brittonii Porter V. comosa Richter V. glandifera Pennell V. micromera Wooton & Standley
Treatment appears in FNA Volume 17. Treatment on page 313. Mentioned on page 306, 314.

Annuals or perennials. Stems erect or prostrate basally, (20–) 30–100 (–170) cm, usually thick-fleshy, glabrous. Leaves: petiole 0–2 (–8) mm (basal lateral branches usually distinctly petiolate to 8 mm) proximally, 0 mm distally; blade (of proximal leaves) ovate, elliptic, or oblong, (15–) 30–80 (–145) × (7–) 10–30 (–45) mm, 1.5–3 times as long as wide, base obtuse proximally to cordate-amplexicaul in middle and ± cuneate distally, margins ± serrulate or denticulate, apex acute, surfaces glabrous, rarely glandular-hairy. Racemes 6–25, axillary, 50–100 (–150) mm, 5–10 mm diam., (20–) 30–40 (–60) -flowered, axis glabrous, rarely glandular-hairy; bracts linear to lanceolate, 2–5 mm, apex acute. Pedicels suberect or arcuate-erect to subpatent, curved upwards in fruit, 3–7 (–10) mm, longer than subtending bracts, glabrous or hairy. Flowers: calyx lobes 2–3 mm, apex acute, glabrous, rarely hairy; corolla lavender to pale blue, rarely pale lilac, (4–) 6–8 mm diam.; stamens 3–3.5 mm; style 1.5–2.5 mm. Capsules slightly compressed in cross-section, globular, (2.5–) 3–3.5 (–4) × 2.5–3.2 (–4) mm, apex rounded or ± emarginate, glabrous or sparsely short-glandular-hairy. Seeds 40–77, yellowbrown, ellipsoid to subglobular, planoconvex, (0.3–) 0.5–0.7 × 0.3–0.5 mm, 0.2–0.3 mm thick, smooth. 2n = 36.


Phenology: Flowering Mar–Sep(–Nov).
Habitat: Stream margins, ditches, banks, springs, swamps, wet meadows.
Elevation: 0–4000 m.

Distribution

Introduced; Alta., B.C., Ont., Ariz., Calif., Colo., Conn., Del., D.C., Fla., Ga., Ill., Ind., Iowa, Kans., Ky., Maine, Mich., Mo., Mont., Nebr., Nev., N.J., N.Mex., N.Y., N.C., N.Dak., Ohio, Okla., Oreg., Pa., R.I., S.Dak., Tenn., Tex., Utah, Va., Wash., W.Va., Wis., Wyo., Eurasia, Africa, Atlantic Islands, likely also in Mexico, Central America, South America

Discussion

Veronica anagallis-aquatica is widespread. It is not clear whether it is native to the flora area; it is certainly introduced in some states and commonly dispersed by human activity. The species varies with water availability. It is frequently confused with V. catenata. Ecological differences (see discussion under 15. V. catenata) may suggest that V. anagallis-aquatica does not occur in most parts of Canada and Alaska from which the species is reported but is present in some states excluded from the distribution area due to the lack of herbarium specimens seen (for example, Alabama, Arkansas, Idaho, Louisiana, Maryland, Massachusetts, Mississippi, South Carolina, and Vermont). Hybrids are frequent in Europe (V. ×lackschewitzkii J. Keller) and have been reported for California and Nebraska (L. R. Heckard and P. Rubtzoff 1977). The sterile hybrids never form capsules and bear relatively long inflorescences. Another closely related species, V. anagalloides Gussone (= V. salina Schur), is not present in North America.

Selected References

None.

Lower Taxa

None.
short-glandular-hairy +, glabrous +  and hairy +
emarginate +, rounded +  and acute +
Dirk C. Albach +
Linnaeus +
glandular-hairy +  and glabrous +
alternate +, opposite +, whorled +, helical +  and subopposite +
cordate-amplexicaul +
cuneate;obtuse +
8 cm80 mm <br />0.08 m <br /> (14.5 cm145 mm <br />0.145 m <br />) +
3 cm30 mm <br />0.03 m <br /> (4.5 cm45 mm <br />0.045 m <br />) +
3 cm30 mm <br />0.03 m <br /> (8 cm80 mm <br />0.08 m <br />) +
oblong;elliptic;oblong;elliptic;ovate +
not leathery +  and not fleshy +
1 cm10 mm <br />0.01 m <br /> (3 cm30 mm <br />0.03 m <br />) +
linear;lanceolate +
0.2 cm2 mm <br />0.002 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
rotate +  and campanulate +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
0.32 cm3.2 mm <br />0.0032 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
globular +  and compressed +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.32 cm3.2 mm <br />0.0032 m <br />) +
darker +, reddish +  and lighter +
Water speedwell +  and véronique mouron-d'eau +
pale lilac +, lavender +  and pale blue +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
campanulate +, short-tubular +  and rotate +
Alta. +, B.C. +, Ont. +, Ariz. +, Calif. +, Colo. +, Conn. +, Del. +, D.C. +, Fla. +, Ga. +, Ill. +, Ind. +, Iowa +, Kans. +, Ky. +, Maine +, Mich. +, Mo. +, Mont. +, Nebr. +, Nev. +, N.J. +, N.Mex. +, N.Y. +, N.C. +, N.Dak. +, Ohio +, Okla. +, Oreg. +, Pa. +, R.I. +, S.Dak. +, Tenn. +, Tex. +, Utah +, Va. +, Wash. +, W.Va. +, Wis. +, Wyo. +, Eurasia +, Africa +, Atlantic Islands +, likely also in Mexico +, Central America +  and South America +
drupe-like +
0–4000 m. +
curved +  and straight +
1 +  and 4 +
Stream margins, ditches, banks, springs, swamps, wet meadows. +
axillary +  and terminal +
alternate +  and opposite +
persistent +  and deciduous +
linear;suborbiculate or triangular-cordate +
serrate +, dentate +  and entire +
denticulate +  and serrulate +
axile +  and parietal +
basal +, apical +  and superior +
tenuinucellate +, unitegmic +  and hemitropous +
campylotropous +, hemianatropous +  and anatropous +
0.7 cm7 mm <br />0.007 m <br /> (1 cm10 mm <br />0.01 m <br />) +
arcuate-erect +  and subpatent +
hairy +  and glabrous +
0.3 cm3 mm <br />0.003 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
5 +  and 4 +
0.2 cm2 mm <br />0.002 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
0 cm0 mm <br />0 m <br /> (0 cm0 mm <br />0 m <br />) +
Flowering Mar–Sep(–Nov). +
(20-)30-40(-60)-flowered +
10 cm100 mm <br />0.1 m <br /> (15 cm150 mm <br />0.15 m <br />) +
5 cm50 mm <br />0.05 m <br /> (10 cm100 mm <br />0.1 m <br />) +
minute +
yellowbrown +
planoconvex +, ellipsoid +  and subglobular +
0.2mm +  and 0.3mm +
Introduced +
0.3 cm3 mm <br />0.003 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
eglandular +  and glandular +
100 cm1,000 mm <br />1 m <br /> (170 cm1,700 mm <br />1.7 m <br />) +
climbing +  and scrambling +
sprawling +, creeping +  and prostrate +
prostrate;erect +
30 cm300 mm <br />0.3 m <br /> (100 cm1,000 mm <br />1 m <br />) +
thick-fleshy +
pointed +  and capitate +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
glandular-hairy +  and glabrous +
Veronica anagallis-aquatica var. terrea +, V. brittonii +, V. comosa +, V. glandifera +  and V. micromera +
Veronica anagallis-aquatica +
Veronica +
species +
gibbous +  and not spurred +
perennial +  and annual +
annual +, biennial +  and perennial +
subshrub +  and herb +