Veronica wormskjoldii

Roemer & Schultes in J. J. Roemer et al.

Syst. Veg. 1: 101. 1817.

Common names: American alpine speedwell
IllustratedEndemic
Synonyms: Veronica alpina subsp. wormskjoldii (Roemer & Schultes) Elenevsky V. alpina var. wormskjoldii (Roemer & Schultes) Hooker
Treatment appears in FNA Volume 17. Treatment on page 309. Mentioned on page 302, 307, 310.

Perennials. Stems erect or ascending, light grayish green distally, unbranched, (3–) 8–50 cm, sparsely to densely villous-hirsute, sometimes also glandular-hairy. Leaves: blade elliptic to lanceolate or oblong-ovate, 8–40 × 5–20 mm, base cuneate, margins entire, dentate, or serrate, apex short-acuminate, surfaces sparsely to densely villous-hirsute or glabrous. Racemes 1, terminal, 5–40 mm, to 60 (–150) mm in fruit, (2–) 5–25-flowered, axis densely villous-hirsute and ± glandular-hairy; bracts linear to lanceolate, 1–8 mm. Pedicels erect, 2–10 (–15) mm, ca. equal to subtending bract, densely villous-hirsute or glandular-hairy. Flowers: calyx lobes (2.5–) 3–5.5 (–7) mm, apex obtuse or acuminate, glabrous or densely glandular-hairy; corolla deep blue-violet or violet-purple to deep blue, rotate, 3–11 mm diam.; stamens 1–2 mm; style 0.8–4 (–6) mm. Capsules compressed in cross-section, oblong-obovoid, 4–6 (–8) × (2.8–) 4–5.5 mm, ca. as long as wide, apex emarginate, usually densely glandular-hairy. Seeds 10–50, straw colored, ellipsoid, flat, 0.7–1 × 0.4–1 mm, 0.05–1 mm thick, very minutely striate.

Distribution

Alta., B.C., N.W.T., Nunavut, Ont., Que., Yukon, Alaska, Ariz., Calif., Colo., Idaho, Maine, Mont., N.H., N.Mex., Nev., Oreg., Utah, Wash., Wyo.

Discussion

Subspecies 2 (2 in the flora).

Veronica wormskjoldii occurs in two cytotypes, a diploid (2n = 18) in western North America and a tetraploid (2n = 36) in eastern North America and Greenland. As demonstrated by D. C. Albach et al. (2006), the tetraploid is likely to be a hybrid of the diploid V. alpina and V. wormskjoldii. In that publication, species rank was used for simplicity, not as a taxonomic conclusion. However, subsequent morphological analyses in conjunction with preparing this treatment revealed that the large variation in the diploid V. wormskjoldii, also supported by the large number of varieties established within that taxon, make it nearly impossible to differentiate the two North American taxa morphologically. In cases where two taxa are morphologically undifferentiable, but karyologically and genetically distinct and geographically clearly separate, the rank of subspecies seems more appropriate.

Selected References

None.

Key

1 Pedicels 2–5 mm; e North America. Veronica wormskjoldii subsp. wormskjoldii
1 Pedicels (2–)5–10(–15) mm; w North America. Veronica wormskjoldii subsp. nutans
... more about "Veronica wormskjoldii"
glandular-hairy +  and glabrous +
Dirk C. Albach +
Roemer & Schultes in J. J. Roemer et al. +
glandular-hairy +  and villous-hirsute +
alternate +, opposite +, whorled +, helical +  and subopposite +
cuneate +
0.8 cm8 mm <br />0.008 m <br /> (4 cm40 mm <br />0.04 m <br />) +
elliptic;lanceolate or oblong-ovate +
? (?) +  and ? (?) +
5mm;20mm +
linear;lanceolate +
0.55 cm5.5 mm <br />0.0055 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.55 cm5.5 mm <br />0.0055 m <br />) +
rotate +  and campanulate +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
0.28 cm2.8 mm <br />0.0028 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
oblong-obovoid +  and compressed +
0.4 cm4 mm <br />0.004 m <br /> (0.55 cm5.5 mm <br />0.0055 m <br />) +
darker +, reddish +  and lighter +
American alpine speedwell +
violet-purple +  and deep blue +
0.3 cm3 mm <br />0.003 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
Alta. +, B.C. +, N.W.T. +, Nunavut +, Ont. +, Que. +, Yukon +, Alaska +, Ariz. +, Calif. +, Colo. +, Idaho +, Maine +, Mont. +, N.H. +, N.Mex. +, Nev. +, Oreg. +, Utah +, Wash. +  and Wyo. +
drupe-like +
curved +  and straight +
1 +  and 4 +
axillary +  and terminal +
alternate +  and opposite +
persistent +  and deciduous +
linear;suborbiculate or triangular-cordate +
serrate +, dentate +  and entire +
palmatifid +, pinnatifid +  and lobed +
axile +  and parietal +
basal +, apical +  and superior +
tenuinucellate +, unitegmic +  and hemitropous +
campylotropous +, hemianatropous +  and anatropous +
1 cm10 mm <br />0.01 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
glandular-hairy +  and villous-hirsute +
0.2 cm2 mm <br />0.002 m <br /> (1 cm10 mm <br />0.01 m <br />) +
5 +  and 4 +
(2-)5-25-flowered +
6 cm60 mm <br />0.06 m <br /> (15 cm150 mm <br />0.15 m <br />) +
0 cm0 mm <br />0 m <br /> (6 cm60 mm <br />0.06 m <br />) +
minute +
ellipsoid +
0.05mm +  and 1mm +
Illustrated +  and Endemic +
green +  and light grayish +
climbing +  and scrambling +
sprawling +, creeping +  and prostrate +
ascending;erect +
8 cm80 mm <br />0.08 m <br /> (50 cm500 mm <br />0.5 m <br />) +
pointed +  and capitate +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
glabrous +  and villous-hirsute +
Veronica alpina subsp. wormskjoldii +  and V. alpina var. wormskjoldii +
Veronica wormskjoldii +
Veronica +
species +
gibbous +  and not spurred +
annual +, biennial +  and perennial +
subshrub +  and herb +