Woodsia scopulina subsp. laurentiana

Windham

Contr. Univ. Michigan Herb. 19: 59. 1993.

Treatment appears in FNA Volume 2.

Scales of stems and petiole bases (at least some) with clusters of dark, occluded cells near center forming narrow, usually discontinuous stripe, scales ovatelanceolate. Pinnae with longest hairs composed of 2–5 cells. Indusial segments narrow, often filamentous distally. Spores averaging 50–57 µm. 2n = 152.


Phenology: Sporulating summer–fall.
Habitat: Cliffs and rocky slopes, found on a variety of substrates including both granite and limestone
Elevation: 0–3000 m

Distribution

V2 314-distribution-map.gif

Alta., B.C., Ont., Que., Ariz., Calif., Colo., Idaho, Mont., Nev., Oreg., S.Dak., Utah, Wash., Wyo.

Discussion

In addition to hybridizing with Woodsia scopulina subsp. scopulina (see comments above), subsp. laurentiana may have crossed with Woodsia oregana subsp. cathcartiana to form W. × maxonii R. M. Tryon. With very few collections and no biosystematic data available, however, the origin of this putative hybrid remains in doubt. Contrary to previous hypotheses (D. F. M. Brown 1964; D. B. Lellinger 1985), Great Lakes populations of W. scopulina were not involved in the origin of the local hybrid known as W. × abbeae (F. S. Wagner 1987).

Selected References

None.

Lower Taxa

None.

"thick" is not a number.

Acute (?) +  and Rounded (?) +
Michael D. Windham +
Windham +
swollen +
simple to commonly +
lanceolate;linear-lanceolate +
reduced +
roundish +
2 (?) +  and 5 (?) +
Alta. +, B.C. +, Ont. +, Que. +, Ariz. +, Calif. +, Colo. +, Idaho +, Mont. +, Nev. +, Oreg. +, S.Dak. +, Utah +, Wash. +  and Wyo. +
0–3000 m +
often +  and glabrous +
Cliffs and rocky slopes, found on a variety of substrates including both granite and limestone +
multicellular +
flattened +
uniseriate +  and multiseriate +
dissected +
monomorphic +
9cm;35cm +
1 cm10 mm <br />0.01 m <br /> (8 cm80 mm <br />0.08 m <br />) +
not articulate +
usually reddish-brown +  and dark purple +
1/5 length of blade;3/4 length of blade +
Sporulating summer–fall. +
not articulate +
longer than wide +
Contr. Univ. Michigan Herb. +
round +, cuplike +, hoodlike +, reniform +, falcate +  and linear +
2 (?) +  and 3 (?) +
bicolored;brown +
persistent +
ovatelanceolate +
not spiny;entire;dentate +
filamentous +
discrete +
cristate +
dictyostelic +, unbranched +  and branched +
erect +  and ascending +
Woodsia obtusa var. lyallii +  and Woodsia oregana var. lyallii +
Woodsia scopulina subsp. laurentiana +
Woodsia scopulina +
subspecies +
oblong +  and round +
on rock +  and terrestrial +
epiphytic +  and hemiepiphytic +
41 +, 39 +  and 38 +