View source for Bucklandiella lawtoniae

{{Treatment/ID
|accepted_name=Bucklandiella lawtoniae
|accepted_authority=(Ireland) Bednarek-Ochyra & Ochyra in R. Ochyra et al.
|publications={{Treatment/Publication
|title=in R. Ochyra et al., Cens. Cat. Polish Mosses,
|place=145. 2003 (as lawtonae),
}}
|basionyms={{Treatment/ID/Basionym
|name=Racomitrium lawtoniae
|authority=Ireland
|rank=species
|publication_title=Bryologist
|publication_place=73: 707, figs. 1–12. 1971 (as lawtonae)
}}
|synonyms=
|hierarchy=Grimmiaceae;Grimmiaceae subfam. Racomitrioideae;Bucklandiella;Bucklandiella sect. Lawtonia;Bucklandiella lawtoniae
|hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Grimmiaceae]]</div><div class="higher-taxon"><small>subfamily</small>[[Grimmiaceae subfam. Racomitrioideae]]</div><div class="higher-taxon"><small>genus</small>[[Bucklandiella]]</div><div class="higher-taxon"><small>section</small>[[Bucklandiella sect. Lawtonia]]</div><div class="higher-taxon"><small>species</small>[[Bucklandiella lawtoniae]]</div></div>
|volume=Volume 27
|mention_page=page 270, 284
|treatment_page=page 279
}}<!--

--><span class="statement" id="st-d0_s0" data-properties="whole_organism fragility;whole_organism size;whole_organism arrangement or pubescence;whole_organism relief;whole_organism texture;whole_organism reflectance;whole_organism coloration;whole_organism coloration;whole_organism coloration;whole_organism coloration;whole_organism coloration;whole_organism growth form"><b>Plants </b>rather robust or less often moderately sized, loosely tufted, coarse and rigid, slightly glistening, yellow, yellow-green, olivaceous, sometimes grayish, in the upper part, brown proximally.</span> <span class="statement" id="st-d0_s1" data-properties="stem atypical some measurement;stem some measurement;stem orientation;stem architecture;stem architecture;stem architecture"><b>Stems </b>5–10 (–15) cm, curved-ascending to erect, simple or sparingly, irregularly branched.</span> <span class="statement" id="st-d0_s2" data-properties="leaf fixation or orientation;leaf arrangement;leaf orientation;leaf course;leaf course;leaf shape or vernation;leaf atypical length;leaf atypical length;leaf length;leaf atypical width;leaf atypical width;leaf width"><b>Leaves </b>erect-appressed to falcate-secund when dry, erect-spreading when wet, straight or curved, sometimes almost circinate, (3.5–) 4–5.5 (–6) × (0.5–) 0.6–0.8 (–0.9) mm;</span> <span class="statement" id="st-d0_s3" data-properties="side quantity;patch architecture">margins broadly recurved on one side and more narrowly on the other side to 1/3–1/2 (–3/4) of the leaf length, 1-stratose throughout or seldom with some bistratose patches distally;</span> <span class="statement" id="st-d0_s4" data-properties="margin orientation;margin length;margin pubescence;awn coloration;awn course;awn orientation;awn atypical some measurement;awn some measurement;awn presence;awn shape;awn shape;awn shape;cell areolation fusion">pilose, awns hyaline, not flexuose, erect-recurved to squarrose, 0.4–1.5 (–2.6) mm, rarely almost absent, flattened, with a distinct cell areolation, remotely sharply denticulate at the margins, distinctly decurrent;</span> <span class="statement" id="st-d0_s5" data-properties="costa architecture;costa shape;costa shape;costa width;costa architecture;median and proximal portion width;median and proximal portion width;adaxial cell atypical quantity;adaxial cell quantity;adaxial cell size;adaxial cell architecture;adaxial cell atypical quantity;adaxial cell quantity;adaxial cell size">costa percurrent, strongly convex abaxially, reniform in the median and proximal portions, 75–100 (–115) µm wide basally, 40–50 µm wide distally, 3 (–5) -stratose basally with (3–) 4–5 enlarged adaxial cells, 2 (–3) -stratose distally, with 2–3 (–4) enlarged adaxial cells;</span> <span class="statement" id="st-d0_s6" data-properties="laminal cell architecture;laminal cell architecture or pubescence or relief">laminal cells 1-stratose throughout, smooth;</span> <span class="statement" id="st-d0_s7" data-properties="lateral wall size;lateral wall shape">basal laminal cells elongate, 25–55 × 9–11 µm, with incrassate, sinuose-nodulose lateral walls;</span> <span class="statement" id="st-d0_s8" data-properties="basal laminal cell shape;basal laminal cell length;basal laminal cell width">alar cells not or slightly differentiated, sometimes slightly yellowish;</span> <span class="statement" id="st-d0_s9" data-properties="cell variability;cell coloration;cell variability;basal marginal border fusion">supra-alar cells hardly differentiated, not forming a distinct basal marginal border;</span> <span class="statement" id="st-d0_s10" data-properties="medial and distal cell shape;medial and distal cell length;medial and distal cell width;inner perichaetial leaf atypical quantity;inner perichaetial leaf shape;inner perichaetial leaf arrangement or shape;inner perichaetial leaf coloration;inner perichaetial leaf shape;inner perichaetial leaf pubescence;distal cell architecture">medial and distal cells rectangular, 12–35 × 5–7 µm. <b>Inner</b> perichaetial leaves (2–3) broadly ovate, convolute, yellowish haline, shortly obtusely pointed or pilose, with thick-walled, chlorophyllose distal cells.</span> <span class="statement" id="st-d0_s11" data-properties="seta some measurement;seta course;seta coloration;seta architecture"><b>Seta </b>4.5–7.5 mm, straight, brown, twisted clockwise.</span> <span class="statement" id="st-d0_s12" data-properties="capsule shape;capsule atypical length;capsule length;capsule width"><b>Capsule </b>obloid to short-cylindric, (1.2–) 1.5–2.5 × 0.6–0.8 mm;</span> <span class="statement" id="st-d0_s13" data-properties="operculum shape;operculum course;operculum some measurement">operculum rostrate, straight, 1–1.5 mm;</span> <span class="statement" id="st-d0_s14" data-properties="peristome tooth some measurement;peristome tooth coloration;peristome tooth architecture or shape;peristome tooth relief;peristome tooth orientation">peristome teeth 250–480 µm, brown, deeply cleft into to branches, strongly papillose, erect.</span> <span class="statement" id="st-d0_s15" data-properties="spore some measurement;spore pubescence or texture"><b>Spores </b>12–17 µm, granulose.</span><!--

-->{{Treatment/Body
|habitat=Humid or wet, mostly shaded acidic rocks, boulders, cliffs and rock outcrops near streams and waterfalls
|elevation=low to moderate elevations (0-800 m)
|distribution=B.C.;Alaska;Wash.
|discussion=<p>Bucklandiella lawtoniae occurs along the coastal region from Kodiak Island and southern Alaska southwards to the Cascade Range in central Washington, with a highly isolated site on Attu Island in the Aleutians. The largest species of the genus, it can be recognized by its characteristic, long and not flexuose, erect-recurved to squarrose, hyaline awns that are distinctly decurrent. Some elongate and pilose plants of B. affinis may imitate B. lawtoniae but that species, apart from a number of differences in structural characters including the hyaline innermost perichaetial leaves, has erect-flexuose awns and leaf margins recurved on both sides towards the apex. The same differences exist between B. lawtoniae and B. heterosticha, in which, additionally, the costa is usually broad, 2-stratose, and canaliculate in the middle and distal parts of the leaf. Additional diagnostic characters of B. lawtoniae are undifferentiated alar cells and the absence of both a basal marginal border and a basal membrane in the peristome. The species appears to be strongly isolated in the genus and on account of its peculiar innermost perichaetial leaves it is positioned in a separate section.</p>
|tables=
|references={{Treatment/Reference
|id=ireland1976a
|text=Ireland, R. R. 1976. Alar cells of Rhacomitrium lawtonae. Bryologist 79: 238–241.
}}
}}<!--

--><!--

-->{{#Taxon:
name=Bucklandiella lawtoniae
|authority=(Ireland) Bednarek-Ochyra & Ochyra in R. Ochyra et al.
|rank=species
|parent rank=section
|synonyms=
|basionyms=Racomitrium lawtoniae
|family=Grimmiaceae
|habitat=Humid or wet, mostly shaded acidic rocks, boulders, cliffs and rock outcrops near streams and waterfalls
|elevation=low to moderate elevations (0-800 m)
|distribution=B.C.;Alaska;Wash.
|reference=ireland1976a
|publication title=in R. Ochyra et al., Cens. Cat. Polish Mosses,
|publication year=
|special status=
|source xml=https://bibilujan@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/84153f6d59a0a91d69695978a64cee7560374f8e/V27/V27_383.xml
|subfamily=Grimmiaceae subfam. Racomitrioideae
|genus=Bucklandiella
|section=Bucklandiella sect. Lawtonia
|species=Bucklandiella lawtoniae
|adaxial cell architecture=2(-3)-stratose
|adaxial cell atypical quantity=3;4
|adaxial cell quantity=2;3
|adaxial cell size=enlarged;enlarged
|awn atypical some measurement=1.5mm;2.6mm
|awn coloration=hyaline
|awn course=not flexuose
|awn orientation=erect-recurved;squarrose
|awn presence=absent
|awn shape=decurrent;denticulate;flattened
|awn some measurement=0.4mm;1.5mm
|basal laminal cell length=25um;55um
|basal laminal cell shape=elongate
|basal laminal cell width=9um;11um
|basal marginal border fusion=distinct
|capsule atypical length=1.2mm;1.5mm
|capsule length=1.5mm;2.5mm
|capsule shape=obloid;short-cylindric
|capsule width=0.6mm;0.8mm
|cell areolation fusion=distinct
|cell coloration=yellowish
|cell variability=differentiated;differentiated
|costa architecture=3(-5)-stratose;percurrent
|costa shape=reniform;convex
|costa width=40um;50um
|distal cell architecture=thick-walled
|inner perichaetial leaf arrangement or shape=convolute
|inner perichaetial leaf atypical quantity=2;3
|inner perichaetial leaf coloration=yellowish
|inner perichaetial leaf pubescence=pilose
|inner perichaetial leaf shape=pointed;ovate
|laminal cell architecture=1-stratose
|laminal cell architecture or pubescence or relief=smooth
|lateral wall shape=sinuose-nodulose
|lateral wall size=incrassate
|leaf arrangement=falcate-secund
|leaf atypical length=5.5mm;6mm
|leaf atypical width=0.8mm;0.9mm
|leaf course=curved;straight
|leaf fixation or orientation=erect-appressed
|leaf length=4mm;5.5mm
|leaf orientation=erect-spreading
|leaf shape or vernation=circinate
|leaf width=0.6mm;0.8mm
|margin length=1-stratose
|margin orientation=recurved
|margin pubescence=pilose
|medial and distal cell length=12um;35um
|medial and distal cell shape=rectangular
|medial and distal cell width=5um;7um
|median and proximal portion width=75um;100um
|operculum course=straight
|operculum shape=rostrate
|operculum some measurement=1mm;1.5mm
|patch architecture=bistratose
|peristome tooth architecture or shape=cleft
|peristome tooth coloration=brown
|peristome tooth orientation=erect
|peristome tooth relief=papillose
|peristome tooth some measurement=250um;480um
|seta architecture=twisted
|seta coloration=brown
|seta course=straight
|seta some measurement=4.5mm;7.5mm
|side quantity=0;1/3-1/2-3/4
|spore pubescence or texture=granulose
|spore some measurement=12um;17um
|stem architecture=branched;sparingly;simple
|stem atypical some measurement=10cm;15cm
|stem orientation=curved-ascending;erect
|stem some measurement=5cm;10cm
|whole_organism arrangement or pubescence=tufted
|whole_organism coloration=brown;grayish;olivaceous;yellow-green;yellow
|whole_organism fragility=robust
|whole_organism growth form=plant
|whole_organism reflectance=glistening
|whole_organism relief=coarse
|whole_organism size=sized
|whole_organism texture=rigid
}}<!--

-->[[Category:Treatment]][[Category:Bucklandiella sect. Lawtonia]]