Veronica wormskjoldii subsp. nutans

(Bongard) Pennell

Rhodora 23: 15. 1921.

Common names: Western American alpine speedwell
Endemic
Basionym: Veronica nutans Bongard Mem. Acad. Imp. Sci. St.-Petersbourg, Ser. 6, Sci. Math. 2: 157. 1832
Synonyms: V. alpina var. alterniflora Fernald V. alpina var. cascadensis Fernald V. alpina var. geminiflora Fernald V. alpina var. nutans (Bongard) Ledebour V. alpina var. unalaschcensis Chamisso & Schlechtendal V. wormskjoldii subsp. alterniflora (Fernald) Pennell
Treatment appears in FNA Volume 17. Treatment on page 310. Mentioned on page 309, 311.
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Pedicels (2–) 5–10 (–15) mm. 2n = 18.


Phenology: Flowering Jun–Aug.
Habitat: Moist alpine meadows, heathlands, and scree, bogs, fens, stream banks, pond and lake margins.
Elevation: 0–3500 m.

Distribution

Alta., B.C., N.W.T., Nunavut, Yukon, Alaska, Ariz., Calif., Colo., Idaho, Mont., Nev., N.Mex., Oreg., Utah, Wash., Wyo.

Discussion

Hybrids of subsp. nutans with Veronica stelleri show morphological intermediacy. Especially plants on the Alaskan coast may have serrate leaves as in V. stelleri and larger corollas as in V. wormskjoldii. The degree and direction of introgression are unknown.

Selected References

None.

Lower Taxa

None.
glandular-hairy +  and glabrous +
acuminate +, obtuse +  and short-acuminate +
Dirk C. Albach +
(Bongard) Pennell +
glandular-hairy +  and villous-hirsute +
alternate +, opposite +, whorled +, helical +  and subopposite +
cuneate +
Veronica nutans +
0.8 cm8 mm <br />0.008 m <br /> (4 cm40 mm <br />0.04 m <br />) +
elliptic;lanceolate or oblong-ovate +
not leathery +  and not fleshy +
0.5 cm5 mm <br />0.005 m <br /> (2 cm20 mm <br />0.02 m <br />) +
linear;lanceolate +
0.1 cm1 mm <br />0.001 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
0.55 cm5.5 mm <br />0.0055 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.55 cm5.5 mm <br />0.0055 m <br />) +
rotate +  and campanulate +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
0.28 cm2.8 mm <br />0.0028 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
oblong-obovoid +  and compressed +
0.4 cm4 mm <br />0.004 m <br /> (0.55 cm5.5 mm <br />0.0055 m <br />) +
darker +, reddish +  and lighter +
Western American alpine speedwell +
violet-purple +  and deep blue +
0.3 cm3 mm <br />0.003 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
Alta. +, B.C. +, N.W.T. +, Nunavut +, Yukon +, Alaska +, Ariz. +, Calif. +, Colo. +, Idaho +, Mont. +, Nev. +, N.Mex. +, Oreg. +, Utah +, Wash. +  and Wyo. +
drupe-like +
0–3500 m. +
curved +  and straight +
1 +  and 4 +
Moist alpine meadows, heathlands, and scree, bogs, fens, stream banks, pond and lake margins. +
axillary +  and terminal +
alternate +  and opposite +
persistent +  and deciduous +
linear;suborbiculate or triangular-cordate +
serrate +, dentate +  and entire +
palmatifid +, pinnatifid +  and lobed +
axile +  and parietal +
basal +, apical +  and superior +
tenuinucellate +, unitegmic +  and hemitropous +
campylotropous +, hemianatropous +  and anatropous +
1 cm10 mm <br />0.01 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
glandular-hairy +  and villous-hirsute +
0.5 cm5 mm <br />0.005 m <br /> (1 cm10 mm <br />0.01 m <br />) +
5 +  and 4 +
Flowering Jun–Aug. +
(2-)5-25-flowered +
6 cm60 mm <br />0.06 m <br /> (15 cm150 mm <br />0.15 m <br />) +
0 cm0 mm <br />0 m <br /> (6 cm60 mm <br />0.06 m <br />) +
minute +
straw colored +
ellipsoid +
0.05mm +  and 1mm +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
3 cm30 mm <br />0.03 m <br /> (8 cm80 mm <br />0.08 m <br />) +
green +  and light grayish +
climbing +  and scrambling +
sprawling +, creeping +  and prostrate +
ascending;erect +
glandular-hairy +  and villous-hirsute +
8 cm80 mm <br />0.08 m <br /> (50 cm500 mm <br />0.5 m <br />) +
pointed +  and capitate +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
glabrous +  and villous-hirsute +
V. alpina var. alterniflora +, V. alpina var. cascadensis +, V. alpina var. geminiflora +, V. alpina var. nutans +, V. alpina var. unalaschcensis +  and V. wormskjoldii subsp. alterniflora +
Veronica wormskjoldii subsp. nutans +
Veronica wormskjoldii +
subspecies +
gibbous +  and not spurred +
annual +, biennial +  and perennial +
subshrub +  and herb +