Ludwigia glandulosa subsp. brachycarpac.

I. Peng

PhytoKeys 145: 58. 2020.

Treatment appears in FNA Volume 10.
Revision as of 20:01, 7 June 2022 by imported>Volume Importer
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Stems rarely reddish green, 10–55 (–90) cm. Leaves: petiole 0–1 cm, blades linear-elliptic to linear, sometimes very narrowly elliptic, those on main axis 3–5 (–7) × 0.3–0.5 (–1) cm, those on branches 0.8–3.6 × 0.2–0.3 (–0.8) cm. Inflorescences: bracteoles attached at base of ovary, 0.4–0.8 × 0.1–0.2 mm. Flowers: sepals 1.1–1.9 × 1–1.8 mm, apex acute or short-acuminate; nectary disc obscurely, minutely papillose; style 0.4–0.8 mm, stigma 0.2–0.3 mm diam. Capsules obscurely 4-angled, 2–5 × 1.3–2 mm, pedicel 0–0.2 mm. Seeds 0.6–0.8 × 0.3–0.4 mm, surface cells elongate transversely to seed length.


Phenology: Flowering Apr–Nov.
Habitat: Ditches, low meadows, coastal prairies, seeps in sandy woods, moist sinkholes in granite outcrops, old clay fields.
Elevation: 0–200 m.

Discussion

Subspecies brachycarpa grows along the Gulf Coast from southwestern Louisiana to Nueces County, Texas, and more sporadically northward in eastern Texas to south-central Oklahoma.

Subspecies brachycarpa was published initially by Peng as a new combination based on Ludwigia cylindrica var. brachycarpa Torrey & A. Gray, not realizing that the variety was based on Jussiaea brachycarpa Lamarck, which Peng considered to be a synonym of L. glandulosa subsp. glandulosa. Therefore, subsp. brachycarpa is a new subspecies but was invalid when published by Peng since it was not accompanied by a Latin description in 1986; that situation has since been remedied.

Selected References

None.

Lower Taxa

None.
sparingly +  and glabrous +
3(-5)-aperturate +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
short-acuminate +  and acute +
Peter C. Hoch +
I. Peng +
attenuate +
0 cm0 mm <br />0 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
Isnardia +
indehiscent +  and loculicidal +
linear-elliptic +  and linear +
3.5 cm35 mm <br />0.035 m <br /> (5.5 cm55 mm <br />0.055 m <br />) +
1.3 cm13 mm <br />0.013 m <br /> (2 cm20 mm <br />0.02 m <br />) +
3 cm30 mm <br />0.03 m <br /> (12 cm120 mm <br />0.12 m <br />) +
lobed +, toothed +  and entire +
deltate +, usually linear +  and lanceolate oblong or obovate +
reduced +
0.3 cm3 mm <br />0.003 m <br /> (2.1 cm21 mm <br />0.021 m <br />) +
reduced +
dark red +  and black +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
narrowly lanceolate +  and sublinear +
0.01 cm0.1 mm <br />1.0e-4 m <br /> (0.02 cm0.2 mm <br />2.0e-4 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (3.6 cm36 mm <br />0.036 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
straight +  and slightly curved +
0.2 cm2 mm <br />0.002 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
spreading +  and erect +
0.13 cm1.3 mm <br />0.0013 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
La. +, Okla. +  and Tex. +
0–200 m. +
pointed +
basifixed +  and versatile +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.11 cm1.1 mm <br />0.0011 m <br />) +
long-pedicellate +  and subsessile +
straight +  and slightly curved +
by a terminal pore +  and dehiscent +
spreading +  and erect +
cylindric +  and clavate turbinate obpyramidal or globose +
Ditches, low meadows, coastal prairies, seeps in sandy woods, moist sinkholes in granite outcrops, old clay fields. +
strigillose +
minute +
perennial +
glandular-puberulent +, hirtellous +, villous +  and strigillose +
slender +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
deciduous +
0.8 cm8 mm <br />0.008 m <br /> (4.5 cm45 mm <br />0.045 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (1 cm10 mm <br />0.01 m <br />) +
5 cm50 mm <br />0.05 m <br /> (7 cm70 mm <br />0.07 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (1 cm10 mm <br />0.01 m <br />) +
3 cm30 mm <br />0.03 m <br /> (5 cm50 mm <br />0.05 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
fringed +  and subentire +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.18 cm1.8 mm <br />0.0018 m <br />) +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.04 cm0.4 mm <br />4.0e-4 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
subcylindric +
0.8mm +  and 1.9mm +
numerous +  and 1 +
submerged +
swollen +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
0 cm0 mm <br />0 m <br /> (0.02 cm0.2 mm <br />2.0e-4 m <br />) +
5 +  and 4 +
attenuate +
0 cm0 mm <br />0 m <br /> (1 cm10 mm <br />0.01 m <br />) +
Flowering Apr–Nov. +
paired +, solitary +  and cluster +
ascending +, decumbent +  and erect +
inconspicuous +
expanded +
basal +  and cauline +
clavate +  and globose +
light-brown +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
50 +  and 400 +
pitted +  and smooth +
kidney--shaped +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.04 cm0.4 mm <br />4.0e-4 m <br />) +
light green +
persistent +
post-anthesis +
5 +  and 4 +
ovate-deltate +
0.1 cm1 mm <br />0.001 m <br /> (0.18 cm1.8 mm <br />0.0018 m <br />) +
parenchymatous +
parietal +, axile +  and placentation +
2 +  and 1 +
unequal +  and subequal +
2 times as many as sepals +
aquatic +, amphibious +  and terrestrial +
lenticular +
2 times as many as sepals +
55 cm550 mm <br />0.55 m <br /> (90 cm900 mm <br />0.9 m <br />) +
reddish green +
sprawling +, decumbent +  and prostrate +
strigillose +, often +  and glabrate +
10 cm100 mm <br />0.1 m <br /> (55 cm550 mm <br />0.55 m <br />) +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.03 cm0.3 mm <br />3.0e-4 m <br />) +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
broadly clavate +  and subglobose +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
sessile +  and petiolate +
green +  and dark reddish +
0.015 cm0.15 mm <br />1.5e-4 m <br /> (0.035 cm0.35 mm <br />3.5e-4 m <br />) +
intrapetiolar +
ovate-triangular +
0.005 cm0.05 mm <br />5.0e-5 m <br /> (0.025 cm0.25 mm <br />2.5e-4 m <br />) +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
aquatic +, amphibious +  and terrestrial +
transversely +  and parallel +
glabrous +, glabrate +  and papillose-strigillose +
papillate +
Gen. +, Ludwigia +, Ludwigia sect. Microcarpium +  and Ludwigiantha +
Ludwigia glandulosa subsp. brachycarpac. +
Ludwigia glandulosa +
subspecies +
3(-5)-aperturate +