Ludwigia lanceolata
Sketch Bot. S. Carolina 1: 213. 1817.
Herbs slender, with well-developed aerenchyma when base submerged, often forming stolons 10–40 cm, 2–3 mm thick, stolons with widely spaced leaves. Stems erect, subterete or slightly ridged, well branched distally, 45–100 cm, glabrous, with raisedlines decurrent from leaf-axils. Leaves alternate; stipules ovate to very broadly ovate, 0.2–0.5 ×0.1–0.3 mm, succulent; stolons: petiole winged, 0.2–1 cm, blade orbiculate or elliptic to broadly elliptic, 0.5–2.7 × 0.6–1.2 cm, base attenuate, apex rounded to acute; main-stem: petiole winged, 0.1–0.5 cm, blade elliptic, oblanceolate, or narrowly oblanceolate to sublinear, 2–7.5 × 0.2–0.8 (–1.4) cm, base narrowly cuneate, sometimes attenuate, margins entire with minute hydathodal glands, apex acute to narrowly acute, leaves on side branches much reduced; bracts sublinear, moderately reduced. Inflorescences open spikes, flowers solitary in distal leaf-axils; bracteoles attached in opposite pairs at or slightly distal to base of ovary, ovate-elliptic to very narrowly elliptic, 1.5–4.3 × 0.4–1.4 mm, often with a swollen base, margins minutely papillose, apex subacute. Flowers: sepals ascending, pale green, broadly ovate-deltate, 1.5–2.5 × 1.8–3.3 mm, margins minutely papillose, apex usually acute, rarely acuminate, surfaces glabrous; petals 0; filaments nearly translucent, 1–1.4 mm, base dilated, anthers 0.4–0.6 (–0.8) × 0.5–0.6 mm; pollen shed in tetrads; ovary broadly obovoid or cupshaped, 2.5–3.5 × 2.2–3.2 mm; nectary disc elevated 0.4–0.6 mm on ovary apex, yellowish green, 1.8–2.6 mm diam., 4-lobed, glabrous; style yellowish green, 0.5–0.7 mm, glabrous, stigma yellowish green, broadly capitate to subglobose, 0.3–0.5 × 0.7–1 mm, not exserted beyond anthers. Capsules obpyramidal, sharply 4-angled and 4-winged, wings 0.3–0.7mm wide, 3.5–5 × 2.5–4.5 mm, hard-walled, dehiscent by apical ring, pedicel 0.1–0.5 mm. Seeds light-brown, narrowly oblong with constricted ends, 0.6–0.8 ×0.2–0.3 mm, surface cells nearly isodiametric. 2n = 32.
Phenology: Flowering Jun–Oct.
Habitat: Ditches, low meadows, cypress swamps, moist pinelands, edges of pocosins, sandy peaty soil.
Elevation: 0–100 m.
Distribution
Fla., Ga., N.C., S.C.
Discussion
Ludwigia lanceolata is fairly uncommon, with scattered populations occurring along the Atlantic coast of southern North Carolina, South Carolina, eastern and southern Georgia, and peninsular Florida. It reaches its western limit in the central panhandle of Florida. C. I. Peng (1988, 1989) found this tetraploid species to be interfertile with other tetraploid species in the section, but few natural hybrids are found, perhaps due to persistent autogamy and habitat specialization.
Selected References
None.