Heterotheca stenophylla

(A. Gray) Shinners

Field & Lab. 19: 68. 1951.

Common names: Stiffleaf goldenaster
Endemic
Basionym: Chrysopsis hispida var. stenophylla A. Gray Boston J. Nat. Hist. 6: 223. 1850
Synonyms: Chrysopsis stenophylla (A. Gray) Greene Chrysopsis villosa var. stenophylla (A. Gray) A. Gray
Treatment appears in FNA Volume 20. Treatment on page 246. Mentioned on page 231, 233, 247.
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Perennials 18–46 (–65) cm; taprooted, sometimes spreading by rhizomes, forming additional clumps. Stems 1–50+, ascending to erect (sometimes brown, or reddish to dark-brown, often proximally brittle), often short-branched in distal 1/2, sparsely to densely long-hispid and strigose (hairs often broken off, especially proximally), sometimes becoming moderately hairy and densely stipitate-glandular distally (axillary fascicles of leaves often present). Leaves generally ascending, congested; proximal cauline petiolate, blades linear-oblanceolate, (15–) 24–42 × (2–) 3–6 (–8.5) mm, bases cuneate to attenuate, margins flat, entire, strigoso-ciliate, proximally long-hispido-strigose, apices acute, faces sparsely to moderately hispido-strigose and sparsely to moderately stipitate-glandular, or moderately to densely hispido-strigose and eglandular; distal sessile (bright green to grayish green), blades linear-oblanceolate to narrowly oblanceolate (rarely broader), (9–) 16.5–30 × (1.5–) 2.2–4.3 (–6) mm, usually little reduced distally (often surpassing heads), bases cuneate to attenuate, margins flat, often abundantly long-hispido-strigose along whole, apices acute, sometime mucronulate to white-spinulose, faces sparsely to moderately densely hispido-strigose (2–65 hairs/mm²; hairs often scabrous, ± pustulate), usually eglandular, sometimes sparsely to densely stipitate-glandular. Heads (1–) 2–16 (–30), in corymbiform arrays, branches ascending. Peduncles 2–16 mm, sparsely to densely hispido-strigose, eglandular or sparsely to densely stipitate-glandular; bracts 1–3, proximal linear-oblanceolate to oblanceolate, sometimes little reduced distally, leaflike. Involucres cylindro-turbinate to narrowly campanulate, (4.5–) 5.5–8 (–11) mm. Phyllaries in 5–6 series, unequal (outer 1/5–1/4 length of inner), narrowly triangular-lanceolate to lanceolate, margins scarious, faces sparsely to moderately strigose, sparsely to densely stipitate-glandular. Ray-florets 10–24 (–36); laminae (5–) 7–11.5 (–16.5) × 0.7–1.5 (–2.1) mm. Disc-florets (12–) 23–55 (–70); corollas slightly ampliate, 4.5–7 mm, throats glabrous, lobes 0.4–0.9 mm, lobes sparsely pilose (hairs on lobes 0.1–0.25 mm). Cypselae monomorphic, obconic, compressed, 1.5–2.5 (–3) mm, ribs 6–10 (sometimes brownish), faces sparsely to moderately strigose; pappi off-white, outer of linear scales 0.25–0.5 mm, inner of 25–40 bristles (4–) 5–6.5 (–7.5) mm, longest weakly clavate. 2n = 18, 36.

Distribution

V20-539-distribution-map.gif

Colo., Iowa, Kans., N.Mex., Nebr., Okla., S.Dak., Tex.

Discussion

Varieties 2 (2 in the flora).

Heterotheca stenophylla is divided into two varieties that differ in gland and hair density. Their ranges overlap to a great extent, and intermediates occur throughout the range. Both H. stenophylla and H. canescens often have numerous, overlapping leaves and often bear axillary fascicles of leaves.

Selected References

None.

Key

1 Distal mid stems glabrate to sparsely hispido-strigose, moderately to densely stipitate- glandular; leaf faces sparsely to moderately hispido-strigose, moderately to densely stipitate-glandular (appearing dark to bright green), hair bases often broad (pustulate); phyllary faces usually sparsely strigose and stipitate-glandular Heterotheca stenophylla var. stenophylla
1 Distal mid stems moderately to densely hispido-strigose, eglandular or sparsely glandular; leaf faces moderately to densely hispido-strigose, eglandular or sparsely stipitate-glandular beneath hairs (appearing pale green to gray-green), hairs rarely broad-based, pustulate; phyllary faces usually moderately strigose, eglandular Heterotheca stenophylla var. angustifolia

"[" is not declared as a valid unit of measurement for this property."]" is not declared as a valid unit of measurement for this property.

... more about "Heterotheca stenophylla"
not tailed +, rounded +  and obtuse +
distinct +
mucronulate;white-spinulose +
subequal +
scarious +
usually triangular +  and linear +
usually deltate +  and lanceolate +
indeterminate +  and determinate +
John C. Semple +
(A. Gray) Shinners +
cuneate +  and attenuate +
Chrysopsis hispida var. stenophylla +
compound +  and simple +
0.9 cm9 mm <br />0.009 m <br /> (1.65 cm16.5 mm <br />0.0165 m <br />) +
0.43 cm4.3 mm <br />0.0043 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
1.65 cm16.5 mm <br />0.0165 m <br /> (3 cm30 mm <br />0.03 m <br />) +
linear-oblanceolate;narrowly oblanceolate +
reduced +
0.22 cm2.2 mm <br />0.0022 m <br /> (0.43 cm4.3 mm <br />0.0043 m <br />) +
rugulose +  and muricate +
barbellulate +  and barbellate +
0.65 cm6.5 mm <br />0.0065 m <br /> (0.75 cm7.5 mm <br />0.0075 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (0.65 cm6.5 mm <br />0.0065 m <br />) +
rhizomatous +  and taprooted +
additional +
Stiffleaf goldenaster +
not 2-lipped +  and actinomorphic +
ampliate +
0.45 cm4.5 mm <br />0.0045 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
compressed +  and obconic +
fertile +  and bisexual +
Colo. +, Iowa +, Kans. +, N.Mex. +, Nebr. +, Okla. +, S.Dak. +  and Tex. +
strigose +  and sparsely densely stipitate-glandular +
pistillate +, staminate +  and neuter +
winged +  and beaked +
dispersed +
paniculiform +
in corymbiform arrays +  and singly +
indeterminate +
heterogamous +  and homogamous +
each +  and sessile +
tan;rusty brown +
attenuate +  and clavate +
0.8 cm8 mm <br />0.008 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
cylindro-turbinate;narrowly campanulate +
0.55 cm5.5 mm <br />0.0055 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
1.15 cm11.5 mm <br />0.0115 m <br /> (1.65 cm16.5 mm <br />0.0165 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.21 cm2.1 mm <br />0.0021 m <br />) +
7mm;11.5mm +
0.7mm;1.5mm +
sessile +  and petiolate +
cauline +  and basal +
erect;spreading +
lanceolate +, triangular +  and deltate +
long-hispido-strigose +
2-carpellate +
in 3 +  and 4 series +
stipitate-glandular +  and hispido-strigose +
0.2 cm2 mm <br />0.002 m <br /> (1.6 cm16 mm <br />0.016 m <br />) +
narrowly triangular-lanceolate +  and lanceolate +
ovate-oblanceolate +
linear-oblanceolate +  and oblanceolate +
Field & Lab. +
bearing subulate enations +, hairy +  and bristly +
subulate;convex +
6 +  and 10 +
3 +  and 40 +
linear +  and linear-lanceolate or triangular +
0.025 cm0.25 mm <br />2.5e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
exalbuminous +
short-branched +
ascending +  and erect +
stipitate-glandular +, hairy +, strigose +  and long-hispid +
1 +  and 50 +
appendaged +  and truncate +
papillate +  and smooth +
Chrysopsis stenophylla +  and Chrysopsis villosa var. stenophylla +
Heterotheca stenophylla +
Heterotheca +
species +
46 cm460 mm <br />0.46 m <br /> (65 cm650 mm <br />0.65 m <br />) +
18 cm180 mm <br />0.18 m <br /> (46 cm460 mm <br />0.46 m <br />) +
tree +, shrub +  and subshrub +