Sidalcea oregana subsp. oregana

[E]
Endemic
Synonyms: Sidalcea maxima M. Peck S. nervata A. Nelson S. oregana var. calva C. L. Hitchcock S. oregana var. maxima (M. Peck) C. L. Hitchcock S. oregana var. nevadensis C. L. Hitchcock S. oregana var. procera C. L. Hitchcock
Treatment appears in FNA Volume 6. Treatment on page 349. Mentioned on page 348, 351.
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Plants (0.3–) 0.8–1.5 m, clumps often 20–30 cm diam., with fibrous-woody taproot or caudex. Stems 3–20, branched or not, usually solid, sometimes ± hollow on older, robust plants, base glabrous or stellate-hairy, hairs to 1 mm, sometimes bristly with appressed, infrequently spreading, hairs, stem distally less hairy, sometimes glabrous or glaucous. Leaves: stipules sometimes purple, linear, 4–6 × 0.5–1 mm; petioles 10–20 (–35) cm on basal leaves, proximalmost 3–5 times as long as blades, greatly reduced distally to 1/2 times blade length; blade 5–15 × 5–15 inflorescence; bracts linear, distal simple and undivided to 2-fid, proximal divided to base, 4–5 mm, usually shorter than young flower buds, usually equaling or longer than pedicels, shorter than calyx. Pedicels 2–5 (–10) mm, 3 mm in flower, 5 mm in fruit. Flowers bisexual or, infrequently, unisexual and pistillate, plants gynodioecious; calyx 5–8 mm, to 10 mm in fruit, usually densely, uniformly stellate-puberulent, marginal hairs sometimes longer, sometimes nearly glabrous; petals bright pink to rose-pink, often pale at base, 9–20 mm, pistillate 5–10 mm, bisexual 10–15 mm; staminal column 4–6 mm, hairy; anthers white; stigmas (6 or) 7 or 8. Schizocarps 6–7 mm diam.; mericarps (6 or) 7 or 8, 2.5–3 mm, usually glabrous, sometimes nearly smooth, back usually rugose-roughened, sides reticulate-veined, sometimes rugose-roughened, mucro 0.3–0.7 mm. Seeds 2 mm. 2n = 20, 40, 60.


Phenology: Flowering (May–)Jun–Aug(–Sep).
Habitat: Moist meadows
Elevation: 500–2500 m

Distribution

V6 629-distribution-map.jpg

B.C., Calif., Idaho, Mont., Nev., Oreg., Utah, Wash., Wyo.

Discussion

Subspecies oregana is variable; it intergrades with subsp. spicata and Sidalcea setosa. C. L. Hitchcock (1957) accepted five varieties within the typical subspecies; morphological intergrades exist. A case can be made for recognition of var. calva C. L. Hitchcock, which has been listed as endangered both federally and in Washington, where it is endemic. These plants are generally robust, sparsely appressed-hairy with four-rayed hairs, the leaves are generally nearly glabrous and fleshy-textured, and the calyx lobes are subglabrous and ciliate. Found in the Wenatchee Mountains, an area of high endemism, var. calva does not appear to be much different from other, nearly glabrous populations elsewhere; it appears to be the only variety with a chromosome count of 2n = 60. This treatment does not accept both subspecies and varieties within Sidalcea; therefore, it has been placed here into synonymy with the wide-ranging, variable typical subspecies. Subspecies oregana can generally be distinguished from the other subspecies by its more-open inflorescences that are elongated in fruit, its multistemmed clumps as much as 30 cm in diameter, its generally stellate-hairy to glabrescent stem bases, its generally uniformly stellate-hairy calyces, its somewhat reticulate-roughened mericarps, and its bracts that are generally equal to or shorter than the young flower buds. Subspecies oregana appears to be the source of commonly sold cultivars. It has been listed as sensitive in Montana and as rare in British Columbia.

Selected References

None.

Lower Taxa

None.

"elongating" is not a number."elongated" is not a number.

60 +, 40 +  and 20 +
rounded +
Steven R. Hill +
stellate-hairy +  and glabrous +
cordate +
Sida oregana +
asymmetric +  and symmetric +
10 cm100 mm <br />0.1 m <br /> (15 cm150 mm <br />0.15 m <br />) +
10 cm100 mm <br />0.1 m <br /> (15 cm150 mm <br />0.15 m <br />) +
3 cm30 mm <br />0.03 m <br /> (10 cm100 mm <br />0.1 m <br />) +
reniform-orbiculate +  and cordate +
3 cm30 mm <br />0.03 m <br /> (10 cm100 mm <br />0.1 m <br />) +
0.6 cm6 mm <br />0.006 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
shorter +  and usually shorter +
divided +, undivided +  and 2-fid +
0.4 cm4 mm <br />0.004 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
reticulate-veined +, not leafy-bracted +  and open +
overlapping +
1.5 cm15 mm <br />0.015 m <br /> (10 cm100 mm <br />0.1 m <br />) +
stellate-puberulent +
0 cm0 mm <br />0 m <br /> (1 cm10 mm <br />0.01 m <br />) +
indehiscent +  and loculicidal +
3(-5)-lobed +  and unlobed +
B.C. +, Calif. +, Idaho +, Mont. +, Nev. +, Oreg. +, Utah +, Wash. +  and Wyo. +
500–2500 m +
sparse;copious +
unisexual +, , +  and bisexual +
elongate +
Moist meadows +
stellate +  and forked +
minute +
0 cm0 mm <br />0 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
axillary +  and terminal +
tough-fibrous +
connate +  and distinct +
simple +, stipulate +, sessile +, subsessile +  and petiolate +
distichous +  and alternate +
10 cm100 mm <br />0.1 m <br /> (35 cm350 mm <br />0.35 m <br />) +
3-5 times as long as blades +
7 cm70 mm <br />0.07 m <br /> (10 cm100 mm <br />0.1 m <br />) +
cauline +  and basal +
reticulate-veined +  and smooth +
entire +  and serrate +
rounded;crenate +
wedge--shaped +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
(3-)5-9-lobed +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.07 cm0.7 mm <br />7.0e-4 m <br />) +
3-40-carpellate +
(1-)2-many +
0.5 cm5 mm <br />0.005 m <br /> (1 cm10 mm <br />0.01 m <br />) +
5 cm50 mm <br />0.05 m <br /> (3 cm30 mm <br />0.03 m <br />) +
overlapping +
1.5 cm15 mm <br />0.015 m <br /> (2 cm20 mm <br />0.02 m <br />) +
pale;bright pink;rose-pink +
adnate +  and distinct +
1 cm10 mm <br />0.01 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
20 cm200 mm <br />0.2 m <br /> (35 cm350 mm <br />0.35 m <br />) +
10 cm100 mm <br />0.1 m <br /> (20 cm200 mm <br />0.2 m <br />) +
Flowering (May–)Jun–Aug(–Sep). +
0-1/2 times blade length +  and 3-5 times as long as blades +
Mem. Amer. Acad. Arts, n. s. +
6mm +  and 7mm +
hairy +  and glabrous +
reniform +
2 cm20 mm <br />0.02 m <br /> (?) +
lobed +  and unlobed +
persistent +
reticulate-veined +
rugose-roughened +
sessile +  and subsessile +
0.6 cm6 mm <br />0.006 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
hollow +, solid +, not +  and branched +
glaucous +, glabrous +  and hairy +
8 +  and 7 +
filiform +, linear +  and capitate +
1-2 times number of carpels +
0.6 cm6 mm <br />0.006 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
linear +  and lanceolate +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
unbranched +  and branched +
hairy +  and glabrous +
Sidalcea maxima +, S. nervata +, S. oregana var. calva +, S. oregana var. maxima +, S. oregana var. nevadensis +  and S. oregana var. procera +
fibrous-woody +
Sidalcea oregana subsp. oregana +
Sidalcea oregana +
subspecies +
30 cm300 mm <br />0.3 m <br /> (80 cm800 mm <br />0.8 m <br />) +
not +  and glaucous +