Drymocallis pseudorupestris var. saxicola

Ertter

J. Bot. Res. Inst. Texas 1: 37, figs. 1M–R. 2007.

Common names: Cliff drymocallis or wood beauty
Endemic
Treatment appears in FNA Volume 9. Treatment on page 290. Mentioned on page 288, 289, 291.

Stems 0.6–2.5 dm, base 1–2 (–3) mm diam., short hairs sparse to moderately abundant, sometimes absent. Basal leaves 3–9 (–15) cm, sparsely to densely hairy (hairs to 1.5 mm), usually densely peglike-glandular, not bristly; leaflet pairs (2–) 3 (–4); terminal leaflet broadly obovate-cuneate, 0.8–2 (–4) × 0.7–2 (–3) cm, teeth single or ± double, 3–8 (–12) per side. Pedicels 3–15 (proximal to 20) mm, usually not bristly, short hairs sparse to moderately abundant (sometimes absent). Flowers (2–) 3–12 (–20); hypanthia and sepals not bristly or bristles less than 1 mm; epicalyx bractlets linear-lanceolate to broadly elliptic, 2–5 × 1–1.5 mm; sepals 4–6 (–7) mm; petals ± overlapping, not red-tinged, narrowly to broadly obovate, 4–8 (–9) × 3–6 (–8) mm; filaments 1–2.5 mm; styles usually golden brown, rarely reddish.


Phenology: Flowering May–Aug.
Habitat: Cliffs, ledges, outcrops, ridges, talus slopes, lava beds, other rocky habitats
Elevation: 1000–3400 m

Distribution

V9 467-distribution-map.jpg

Alta., B.C., Calif., Idaho, Mont., Nev., Oreg., Utah, Wash., Wyo.

Discussion

Variety saxicola accommodates the bulk of specimens previously placed in Potentilla glandulosa var. pseudorupestris, minus the extremes at the northeastern and southern ends of the species range. The circumscription here encompasses significant heterogeneity, which might be resolved into additional taxa upon further analysis [for example, the dwarfed high elevation form of P. glandulosa noted by N. H. Holmgren (1997b)]. Some collections from Alberta and Washington have petals as large as those of var. pseudorupestris but are here included in var. saxicola on the basis of stature. Plants from southwestern Idaho and southeastern Oregon combine features of D. lactea and D. pseudorupestris, some being atypically tall (to 5.5 dm) but with the vestiture and saxicolous preference of var. saxicola. In contrast, populations on Steens Mountain, Oregon, including the type of D. pumila Rydberg, have the typical habit of var. saxicola but the vestiture of D. lactea; their optimal placement is unresolved. In California, var. saxicola occurs in the Cascade Range (Mount Lassen, Mount Shasta) and extends sporadically south through the Sierra Nevada to Tulare County, intergrading with var. crumiana and D. lactea.

Selected References

None.

Lower Taxa

None.

"thin" is not a number."wide" is not a number."dm" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property.

light-brown +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
acute;obtuse +
Barbara Ertter +
Ertter +
compound +  and simple +
opposite +  and alternate +
9 cm90 mm <br />0.09 m <br /> (15 cm150 mm <br />0.15 m <br />) +
3 cm30 mm <br />0.03 m <br /> (9 cm90 mm <br />0.09 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
Potentilla pseudorupestris +
oblanceolate-oblong +
1 cm10 mm <br />0.01 m <br /> (25 cm250 mm <br />0.25 m <br />) +
coriaceous +
reduced +
(10-)20-40(-50)° +
0 cm0 mm <br />0 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
free +  and distinct +
Cliff drymocallis or wood beauty +
Alta. +, B.C. +, Calif. +, Idaho +, Mont. +, Nev. +, Oreg. +, Utah +, Wash. +  and Wyo. +
not +  and aggregated +
1000–3400 m +
0.2 cm2 mm <br />0.002 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
linear-lanceolate +  and broadly elliptic +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
3 +  and 12 +
unisexual +  and bisexual +
not +  and aggregated +
deciduous +
less;10;80 +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
Cliffs, ledges, outcrops, ridges, talus slopes, lava beds, other rocky habitats +
sparse +  and moderately abundant +
open +, leafy +  and 2-40-flowered +
1/6 +  and 3/4-4/5 +
crassinucellate +
opposite +  and alternate +
marcescent +
winter-persistent +
hairy +  and glabrate +
overlapping +  and not confluent +
elliptic-ovate +  and flabellate or rhombic +
toothed +, or +, flat +, entire +  and more or less toothed +
inferior +  and superior +
clustered +, biseriate +  and superposed +
0.3 cm3 mm <br />0.003 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
overlapping +
0.8 cm8 mm <br />0.008 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
0 (?) +  and 4 (?) +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
not red-tinged +
free +  and distinct +
obovate +
0.3 cm3 mm <br />0.003 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
Flowering May–Aug. +
adnate +, free +, connate +  and distinct +
J. Bot. Res. Inst. Texas +
not arillate +
0.6 cm6 mm <br />0.006 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
persistent +
free +  and distinct +
not bristly +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
free +  and distinct +
branched +  and simple +
openly tufted +  and loosely spaced +
stramineous +, reddish +  and green +
1 +  and 20 +
palmate +  and pinnate +
narrowly;broadly-triangular-ovate +
reddish +  and golden brown +
deciduous +
distinct +
fusiform +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
Drymocallis glandulosa subsp. pseudorupestris +, P. glandulosa subsp. pseudorupestris +, P. glandulosa var. pseudorupestris +  and P. rupestris var. americana +
Drymocallis pseudorupestris var. saxicola +
Drymocallis pseudorupestris +
variety +
2 cm20 mm <br />0.02 m <br /> (4 cm40 mm <br />0.04 m <br />) +
2 cm20 mm <br />0.02 m <br /> (3 cm30 mm <br />0.03 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (2 cm20 mm <br />0.02 m <br />) +
obovate-cuneate +
0.7 cm7 mm <br />0.007 m <br /> (2 cm20 mm <br />0.02 m <br />) +
horseshoe--shaped +
3 (?) +  and 8 (?) +
inconspicuous +
hemispheric +  and conic +
enlarged +
peglike-glandular +  and septate-glandular +