Horkelia daucifolia var. caruifolia

(Rydberg ex Howell) Ertter & Reveal

Novon 17: 319. 2007.

Common names: Rogue River horkelia
Endemic
Basionym: Horkelia caruifolia Rydberg ex Howell Fl. N.W. Amer., 181. 1898
Synonyms: Potentilla caruifolia (Rydberg ex Howell) Crum P. caruifolia var. divergens Crum
Treatment appears in FNA Volume 9. Treatment on page 267. Mentioned on page 266.

Plants rosette-forming to ± matted. Basal leaves green to grayish, (4–) 5–9 (–15) cm; leaflets 5–10 (–15) mm, lobes 4–15, linear, 0.4–1 mm wide. Pedicels 2–7 (–10) mm, puberulent, eglandular. Flowers: petals broadly obcordate, (3.5–) 5–8 × (4–) 5–8 mm; filaments 2–3 mm.


Phenology: Flowering summer.
Habitat: Grassy openings in mixed oak and conifer woodlands, on non-serpentine soil
Elevation: 400–1100 m

Discussion

Except for a single population in extreme southeastern Klamath County (noted under var. daucifolia), all populations of Horkelia daucifolia from Oregon are here referred to var. caruifolia (B. Ertter and J. L. Reveal 2007). The variety is found in the Rogue River drainage in Jackson and Josephine counties and might prove to be of conservation concern. Collections from near Ashland, including the type of H. caruifolia, are not as distinct from var. daucifolia as are populations from north of the Rogue River Valley, which have particularly broad petals and mat-forming capabilities. These northern populations include the type of Potentilla caruifolia var. divergens, which was described on the basis of its widely open inflorescence; subsequent collections from the same general area (near Trail) have the full range of inflorescence compactness found elsewhere in H. daucifolia.

Selected References

None.

Lower Taxa

None.

"thin" is not a number."dm" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property.

smooth +  and more or less roughened +
2.4mm +  and 3mm +
shorter than wide +  and longer +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
rounded +
Barbara Ertter +  and James L. Reveal +
(Rydberg ex Howell) Ertter & Reveal +
compound +  and simple +
opposite +  and alternate +
12 cm120 mm <br />0.12 m <br /> (17 cm170 mm <br />0.17 m <br />) +
9 cm90 mm <br />0.09 m <br /> (15 cm150 mm <br />0.15 m <br />) +
3 cm30 mm <br />0.03 m <br /> (4 cm40 mm <br />0.04 m <br />) +
green +  and grayish +
5 cm50 mm <br />0.05 m <br /> (12 cm120 mm <br />0.12 m <br />) +
weakly planar;loosely cylindric +
5 cm50 mm <br />0.05 m <br /> (9 cm90 mm <br />0.09 m <br />) +
0.8 cm8 mm <br />0.008 m <br /> (3 cm30 mm <br />0.03 m <br />) +
flattened +
Horkelia caruifolia +
30 cm300 mm <br />0.3 m <br /> (40 cm400 mm <br />0.4 m <br />) +
planar +  and cylindric +
2 cm20 mm <br />0.02 m <br /> (30 cm300 mm <br />0.3 m <br />) +
coriaceous +
reduced +
free +  and distinct +
Rogue River horkelia +
open to congested +
not +  and aggregated +
400–1100 m +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
linear +  and linear-lanceolate +
0.5 cm5 mm <br />0.005 m <br /> (?) +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
flattened +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0.5mm;1.5mm +
in more or less corymbiform clusters +  and arranged +
1 cm10 mm <br />0.01 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
unisexual +  and bisexual +
not +  and aggregated +
obliquely ovoid +  and reniform +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
sparse +  and dense +
noncapitate +  and capitate +
Grassy openings in mixed oak and conifer woodlands, on non-serpentine soil +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
pilose +  and glabrous +
0 +  and 1/2 +
0.1 cm1 mm <br />0.001 m <br /> (0.55 cm5.5 mm <br />0.0055 m <br />) +
crassinucellate +
opposite +  and alternate +
marcescent +
winter-persistent +
1.5 cm15 mm <br />0.015 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
1 cm10 mm <br />0.01 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
divided +, obovate +  and broadly obcordate +
0.5 cm5 mm <br />0.005 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (?) +  and 2 cm20 mm <br />0.02 m <br /> (?) +
linear +  and oblanceolate +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (?) +  and 0.1 cm1 mm <br />0.001 m <br /> (?) +
palmate +  and pinnate +
shallowly toothed +  and deeply divided +
aromatic +
inferior +  and superior +
clustered +, biseriate +  and superposed +
0.7 cm7 mm <br />0.007 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0 (?) +  and 4 (?) +
0.4 cm4 mm <br />0.004 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
yellowish +, white +  and cream +
free +  and distinct +
obcordate +
0.5 cm5 mm <br />0.005 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
Flowering summer. +
adnate +, free +, connate +  and distinct +
not arillate +
linear;filiform +
persistent +
free +  and distinct +
obtuse +  and acute +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.75 cm7.5 mm <br />0.0075 m <br />) +
free +  and distinct +
branched +  and simple +
stramineous +, reddish +  and green +
ascending +  and erect +
1 +  and 10 +
palmate +  and pinnate +
1-3-lobed;3-7-lobed;divided +
0.8 cm8 mm <br />0.008 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
deciduous +
distinct +
subterminal +
filiform +
rough-thickened +
0.2 cm2 mm <br />0.002 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
Potentilla caruifolia +  and P. caruifolia var. divergens +
Horkelia daucifolia var. caruifolia +
Horkelia daucifolia +
variety +
inconspicuous +
enlarged +
green +  and grayish +
plant +, rosette-forming +  and more or less matted +
hairy +  and inconspicuously conspicuously glandular +