Ludwigia arcuata

Walter

Fl. Carol., 89. 1788.

Common names: Piedmont primrose-willow
Synonyms: Isnardia arcuata (Walter) Kuntze I. pedunculosa (Michaux) de Candolle Ludwigia pedunculosa Michaux Ludwigiantha arcuata (Walter) Small
Treatment appears in FNA Volume 10.

Herbs usually creeping and rooting at nodes, forming mats. Stems prostrate or decumbent and ascending at tips, slightly ridged, often well branched, 5–70 cm, glabrate to sparsely strigillose, denser on distal parts. Leaves opposite; stipules narrowly deltate or ovate, 0.05–0.15 × 0.05–0.1 mm; submerged stems: petiole 0–0.2 cm, blade narrowly linear, 1.9–4 ×0.1–0.25 cm; emergent stems: petiole 0–0.2 cm, blade narrowly elliptic or narrowly oblanceolate-elliptic to linear, 0.6–1.8 × 0.2–0.5 cm, base narrowly cuneate, margins entire, apex acute, surfaces glabrous or sparingly strigillose on margins and abaxial midveins; bracts reduced. Inflorescences in racemes or spikes, well-formed on ascending stems, not on prostrate stems; bracteoles attached at base of ovary or 1.5–8 mm proximally on pedicels, sublinear to very narrowly elliptic, 1.4–5 × 0.2–0.8 mm, apex acute, surfaces minutely strigillose. Flowers: sepals reflexed or spreading, green, lanceolate-deltate, 5.2–10 ×1.5–2.7 mm, with 3 prominent parallel veins, margins entire and minutely strigillose, apex acute or elongate-acuminate, surfaces minutely strigillose abaxially; petals rarely caducous, elliptic-obovate to spatulate-obovate, 7–11 × 4.5–8 mm, base attenuate, apex rounded; filaments initially spreading, becoming erect, yellow, 2.5–4.5 mm, anthers 1.3–2 × 0.7–1.1 mm; pollen shed in tight tetrads; ovary cylindric to funnelform, 4–5.5 × 1.5–2.8 mm; nectary disc elevated 0.6–1 mm on ovary apex, 1.5–2.6 mm diam., bright-yellow, with 4 distinct domed lobes, minutely strigillose between lobes or glabrous; style yellow, 2.3–4 (–4.8) mm, glabrous, stigma yellow, broadly capitate, 0.3–0.6 × 0.6–1.8 mm, as long as or exserted beyond anthers. Capsules clavate, subterete, sometimes slightly curved, 5.5–10 × 2.3–4 mm, hard-walled, irregularly dehiscent, pedicel (12–) 17–45 mm. Seeds light to dark-brown, elliptic-oblong, 0.5–0.7 × 0.3–0.4 mm, surface cells transversely elongate. 2n = 32.


Phenology: Flowering Mar–Aug.
Habitat: Roadside ditches, edges of lakes or ponds, swampy prairies, springs, mucky or sandy beach strands.
Elevation: 0–150 m.

Discussion

Ludwigia arcuata is common in its range, but geographically restricted to central and western parts of peninsular Florida and adjacent Georgia, extending to southern South Carolina. Disjunct populations have been collected in Bibb County in central Georgia and Mobile County, Alabama.

The tetraploid Ludwigia arcuata has the largest flowers in sect. Isnardia and is the most consistently outcrossing species; C. I. Peng (1989) reported abun­dant insect visitors on this species. It is morphologically most similar to the hexaploid L. brevipes, with which it shares two genomes (Peng).

Selected References

None.

Lower Taxa

None.
... more about "Ludwigia arcuata"
3(-5)-aperturate +
0.13 cm1.3 mm <br />0.0013 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.11 cm1.1 mm <br />0.0011 m <br />) +
rounded +, elongate-acuminate +  and acute +
Peter C. Hoch +
Walter +
attenuate +  and cuneate +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
Isnardia +
indehiscent +  and loculicidal +
oblanceolate-elliptic to linear +, oblanceolate-elliptic +  and elliptic +
0.6 cm6 mm <br />0.006 m <br /> (1.8 cm18 mm <br />0.018 m <br />) +
lobed +, toothed +  and entire +
deltate +, usually linear +  and lanceolate oblong or obovate +
reduced +
0.2 cm2 mm <br />0.002 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
reduced +
dark red +  and black +
0.14 cm1.4 mm <br />0.0014 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
sublinear to very +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
0.55 cm5.5 mm <br />0.0055 m <br /> (1 cm10 mm <br />0.01 m <br />) +
spreading +  and erect +
subterete +  and clavate +
0.23 cm2.3 mm <br />0.0023 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
Piedmont primrose-willow +
Ala. +, Fla. +, Ga. +  and S.C. +
0–150 m. +
basifixed +  and versatile +
erect +  and spreading +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
long-pedicellate +  and subsessile +
straight +  and slightly curved +
by a terminal pore +  and dehiscent +
spreading +  and erect +
cylindric +  and clavate turbinate obpyramidal or globose +
Roadside ditches, edges of lakes or ponds, swampy prairies, springs, mucky or sandy beach strands. +
perennial +
glandular-puberulent +, hirtellous +, villous +  and strigillose +
deciduous +
distinct +
strigillose +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.26 cm2.6 mm <br />0.0026 m <br />) +
glabrous +  and strigillose +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.55 cm5.5 mm <br />0.0055 m <br />) +
cylindric +
1.5mm +  and 2.8mm +
numerous +  and 1 +
submerged +
swollen +
1.2 cm12 mm <br />0.012 m <br /> (1.7 cm17 mm <br />0.017 m <br />) +
1.7 cm17 mm <br />0.017 m <br /> (4.5 cm45 mm <br />0.045 m <br />) +
7mm +  and 11mm +
5 +  and 4 +
elliptic-obovate +  and spatulate-obovate +
0.45 cm4.5 mm <br />0.0045 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
0 cm0 mm <br />0 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
Flowering Mar–Aug. +
Fl. Carol., +
paired +, solitary +  and cluster +
ascending +, decumbent +  and erect +
inconspicuous +
expanded +
basal +  and cauline +
clavate +  and globose +
light +  and dark-brown +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.07 cm0.7 mm <br />7.0e-4 m <br />) +
50 +  and 400 +
pitted +  and smooth +
elliptic-oblong +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.04 cm0.4 mm <br />4.0e-4 m <br />) +
persistent +
post-anthesis +
spreading +  and reflexed +
5 +  and 4 +
lanceolate-deltate +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.27 cm2.7 mm <br />0.0027 m <br />) +
parenchymatous +
parietal +, axile +  and placentation +
2 +  and 1 +
unequal +  and subequal +
2 times as many as sepals +
aquatic +, amphibious +  and terrestrial +
lenticular +
2 times as many as sepals +
prostrate +  and decumbent +
emergent +  and submerged +
glabrate +  and sparsely strigillose +
5 cm50 mm <br />0.05 m <br /> (70 cm700 mm <br />0.7 m <br />) +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
hemispherical +, capitate +, lobed +  and entire +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.18 cm1.8 mm <br />0.0018 m <br />) +
sessile +  and petiolate +
green +  and dark reddish +
0.005 cm0.05 mm <br />5.0e-5 m <br /> (0.015 cm0.15 mm <br />1.5e-4 m <br />) +
intrapetiolar +
ovate +  and deltate +
0.005 cm0.05 mm <br />5.0e-5 m <br /> (0.01 cm0.1 mm <br />1.0e-4 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.48 cm4.8 mm <br />0.0048 m <br />) +
0.23 cm2.3 mm <br />0.0023 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
aquatic +, amphibious +  and terrestrial +
strigillose +  and glabrous +
papillate +
Isnardia arcuata +, I. pedunculosa +, Ludwigia pedunculosa +  and Ludwigiantha arcuata +
Ludwigia arcuata +
Ludwigia sect. Isnardia +
species +
3(-5)-aperturate +