Ludwigia ravenii

C. I. Peng

Syst. Bot. 9: 129, fig. 1. 1984.

Common names: Raven’s primrose-willow
Treatment appears in FNA Volume 10.

Herbs slender, rarely with aerenchyma when base sub­merged, forming slender, gla­brate stolons 10–18 cm, 0.6–1.5 mm thick, occasionally bearing flowers and fruits. Stems erect, slightly ridged, usually well branched, (15–) 35–90 cm, densely hirtellous. Leaves alter­nate; stipules lanceolate to broadly deltate, 0.2–0.5 × 0.1–0.3 mm; stolons: petiole 0.1–0.3 cm, blade elliptic to orbiculate, 1–1.8 × 0.6–1.4 cm, base attenuate, apex rounded to acute; stems: petiole narrowly winged, 0.1–0.8 cm, blade narrowly lanceolate-elliptic, 1.3–6.5 × 0.4–1.5 cm, base attenuate, margins entire with minute hydathodal glands, apex acute, surfaces densely hirtellous; bracts not much reduced. Inflorescences leafy racemes, flowers solitary in leaf-axils; bracteoles attached near base of ovary, lanceolate or elliptic to narrowly so, (1.5–) 2–4.3 ×0.3–0.9 mm, apex acuminate, surfaces hirtellous. Flowers: sepals ascending-spreading, green, broadly ovate-deltate, 1.5–3 × 1.4–2.1 mm, margins entire, apex acuminate, surfaces densely hirtellous; petals 0; filaments light green, 0.7–1.1 mm, anthers 0.3–0.4 × 0.4–0.5 mm; pollen shed in tetrads; ovary obovoid to obconic, 2.8–3.5 × 2–3 mm; nectary disc elevated 0.3–0.4 mm on ovary apex, light green, 1.4–2.5 mm diam., 4-lobed, glabrous; style light green, 0.3–0.5 mm, glabrous, stigma clavate to subcapitate, 0.4–0.6 × 0.3–0.5 mm, not exserted beyond anthers. Capsules oblong-obovoid, subterete to scarcely 4-angled, (3–) 4–5 (–5.3) × 2.5–3.5 (–4) mm, hard-walled, irregularly dehiscent, pedicel 0.2–0.5 mm. Seeds light-brown, elliptic-oblong with slightly curved ends, 0.5–0.7 ×0.3–0.4 mm, surface cells elongate transversely to seed length. 2n = 32.


Phenology: Flowering Jul–Sep.
Habitat: Wet, peaty habitats, ditches, margins of ponds, bogs, swamps.
Elevation: 0–100 m.

Discussion

Ludwigia ravenii is an uncommon species occur­ring in scattered populations in coastal southeastern Virginia and eastern North Carolina, with single disjunct populations in southeastern South Carolina and northeastern Florida. C. I. Peng (1989) observed its similarity to L. pilosa by virtue of its dense, hirtellous pubescence, but noted its smaller, more consistently autogamous flowers. It is in the Center for Plant Conservation’s National Collection of Endangered Plants.

Selected References

None.

Lower Taxa

None.
... more about "Ludwigia ravenii"
10 cm100 mm <br />0.1 m <br /> (18 cm180 mm <br />0.18 m <br />) +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
3(-5)-aperturate +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.04 cm0.4 mm <br />4.0e-4 m <br />) +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
acuminate +, acute +  and rounded +
Peter C. Hoch +
C. I. Peng +
attenuate +
Isnardia +
indehiscent +  and loculicidal +
lanceolate-elliptic +
1.3 cm13 mm <br />0.013 m <br /> (6.5 cm65 mm <br />0.065 m <br />) +
lobed +, toothed +  and entire +
elliptic +  and orbiculate +
reduced +
0.4 cm4 mm <br />0.004 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
reduced +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
dark red +  and black +
0.2 cm2 mm <br />0.002 m <br /> (0.43 cm4.3 mm <br />0.0043 m <br />) +
elliptic +  and lanceolate +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.09 cm0.9 mm <br />9.0e-4 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (0.53 cm5.3 mm <br />0.0053 m <br />) +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
straight +  and slightly curved +
0.4 cm4 mm <br />0.004 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
spreading +  and erect +
subterete +  and scarcely 4-angled +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
Raven’s primrose-willow +
Fla. +, N.C. +, S.C. +  and Va. +
0–100 m. +
basifixed +  and versatile +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.11 cm1.1 mm <br />0.0011 m <br />) +
long-pedicellate +  and subsessile +
straight +  and slightly curved +
by a terminal pore +  and dehiscent +
spreading +  and erect +
cylindric +  and clavate turbinate obpyramidal or globose +
Wet, peaty habitats, ditches, margins of ponds, bogs, swamps. +
perennial +
glandular-puberulent +, hirtellous +, villous +  and strigillose +
slender +
opposite +  and alternate +
deciduous +
0.14 cm1.4 mm <br />0.0014 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.04 cm0.4 mm <br />4.0e-4 m <br />) +
0.28 cm2.8 mm <br />0.0028 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
obovoid +  and obconic +
2mm +  and 3mm +
numerous +  and 1 +
submerged +
swollen +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
5 +  and 4 +
0.1 cm1 mm <br />0.001 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
Flowering Jul–Sep. +
paired +, solitary +  and cluster +
ascending +, decumbent +  and erect +
inconspicuous +
expanded +
basal +  and cauline +
clavate +  and globose +
light-brown +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.07 cm0.7 mm <br />7.0e-4 m <br />) +
50 +  and 400 +
pitted +  and smooth +
elliptic-oblong +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.04 cm0.4 mm <br />4.0e-4 m <br />) +
persistent +
post-anthesis +
ascending-spreading +
5 +  and 4 +
ovate-deltate +
0.14 cm1.4 mm <br />0.0014 m <br /> (0.21 cm2.1 mm <br />0.0021 m <br />) +
parenchymatous +
parietal +, axile +  and placentation +
2 +  and 1 +
unequal +  and subequal +
2 times as many as sepals +
aquatic +, amphibious +  and terrestrial +
lenticular +
2 times as many as sepals +
15 cm150 mm <br />0.15 m <br /> (35 cm350 mm <br />0.35 m <br />) +
sprawling +, decumbent +  and prostrate +
hirtellous +
35 cm350 mm <br />0.35 m <br /> (90 cm900 mm <br />0.9 m <br />) +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
clavate +  and subcapitate +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
sessile +  and petiolate +
green +  and dark reddish +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
intrapetiolar +
lanceolate +  and broadly deltate +
0.01 cm0.1 mm <br />1.0e-4 m <br /> (0.03 cm0.3 mm <br />3.0e-4 m <br />) +
slender +
light green +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
aquatic +, amphibious +  and terrestrial +
papillate +
Gen. +, Ludwigia +, Ludwigia sect. Microcarpium +  and Ludwigiantha +
Ludwigia ravenii +
Ludwigia sect. Isnardia +
species +
3(-5)-aperturate +