View source for Grimmia hamulosa

{{Treatment/ID
|accepted_name=Grimmia hamulosa
|accepted_authority=Lesquereux
|publications={{Treatment/Publication
|title=Mem. Calif. Acad. Sci.
|place=1: 14. 1868,
}}
|basionyms=
|synonyms={{Treatment/ID/Synonym
|name=Grimmia brevirostris
|authority=R. S. Williams
|rank=species
}}
|hierarchy=Grimmiaceae;Grimmiaceae subfam. Grimmioideae;Grimmia;Grimmia subg. Litoneuron;Grimmia hamulosa
|hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Grimmiaceae]]</div><div class="higher-taxon"><small>subfamily</small>[[Grimmiaceae subfam. Grimmioideae]]</div><div class="higher-taxon"><small>genus</small>[[Grimmia]]</div><div class="higher-taxon"><small>subgenus</small>[[Grimmia subg. Litoneuron]]</div><div class="higher-taxon"><small>species</small>[[Grimmia hamulosa]]</div></div>
|volume=Volume 27
|mention_page=page 228, 244, 544
|treatment_page=page 243
}}<!--

--><span class="statement" id="st-d0_s0" data-properties="whole_organism coloration;whole_organism growth form;patch prominence or shape"><b>Plants </b>in flat patches, blackish green.</span> <span class="statement" id="st-d0_s1" data-properties="stem some measurement"><b>Stems </b>1–1.5 cm.</span> <span class="statement" id="st-d0_s2" data-properties="leaf shape;leaf shape;leaf length or size;base architecture or fixation;base shape;point shape;point shape;point some measurement;margin shape;margin orientation;intermarginal band presence;intermarginal band shape;awn height or length or size;awn coloration;awn presence;awn shape;costa size or width"><b>Leaves </b>homomallous-falcate, oblong-lanceolate, from a clasping base tapering to a long, subulate uncinate point, 2–3.5 mm, margins plane, erect distally, intermarginal bands absent, usually muticous, occasionally short hyaline awns present, not decurrent, costa narrow proximally;</span> <span class="statement" id="st-d0_s3" data-properties="laminal cell shape;laminal cell course;laminal cell architecture;laminal cell architecture">basal juxtacostal laminal cells short-rectangular, straight to sinuose, thick-walled;</span> <span class="statement" id="st-d0_s4" data-properties="basal marginal laminal cell shape;basal marginal laminal cell architecture">basal marginal laminal cells quadrate to short-rectangular, thick-walled;</span> <span class="statement" id="st-d0_s5" data-properties="medial laminal cell shape;medial laminal cell architecture;medial laminal cell architecture">medial laminal cells irregularly rounded to quadrate, sinuose, thick-walled;</span> <span class="statement" id="st-d0_s6" data-properties="distal laminal cell architecture;distal laminal cell shape">distal laminal cells 2-stratose, irregularly rounded to quadrate.</span> <span class="statement" id="st-d0_s7" data-properties="perichaetial leaf size"><b>Perichaetial </b>leaves not enlarged.</span> <span class="statement" id="st-d0_s8" data-properties="seta course;seta some measurement"><b>Seta </b>straight, 2.5–3.5 mm.</span> <span class="statement" id="st-d0_s9" data-properties="capsule presence;capsule position;capsule shape;capsule reflectance;capsule architecture or pubescence or relief;capsule coloration or pubescence or relief;stoma presence;annulus presence;operculum shape;basal segment architecture or pubescence or relief;distal segment relief"><b>Capsule </b>occasionally present, exserted, oblate, shiny, smooth, becoming striate when dry, stomata absent, annulus absent, operculum conical, peristome split in distal half, basal segments smooth, distal segments papillose.</span> <span class="statement" id="st-d0_s10" data-properties=""><b>Calyptra </b>unknown.</span><!--

-->{{Treatment/Body
|habitat=Dry granitic rock and boulders
|elevation=moderate to high elevations (1500-3500 m)
|distribution=Calif.;Oreg.;Wash.
|discussion=<p>Of conservation concern.</p><!--
--><p>Few collections of Grimmia hamulosa exist, its habitat is not well defined, and it is a somewhat confusing species. The original description is incomplete and partly incorrect, e.g. it lacks discussion and comparison with related taxa, and the seta is straight, not bent, as stated in the protologue. The species is characterized by: (1) a glossy habit with long tapering, homomallous, muticous leaves with plane margins; (2) a broad, weakly outlined costa that fills the distal part of the 2-stratose lamina; and (3) a glossy exserted capsule that lacks an annulus. Despite its unique leaf characters and lack of an annulus, Hastings has put it in subg. Litoneuron based on its thick, concave leaves with plane margins, costa not projecting prominently from the lamina, and exserted capsule on a straight seta. In 1999, Greven found G. hamulosa richly fruiting in Yosemite National Park. Although isotypes studied by Greven have muticous leaves, in some of the plants from Yosemite the leaves have inconspicuous short awns. However, these plants do not deviate in other aspects from the type specimen. Small specimens of this species resemble G. fragilis Schimper, a form of G. montana described from southern Europe. These plants are characterized by long, glossy leaves usually with broken tips. In G. hamulosa, the leaf tips are also frequently broken; however, in G. montana the costa is clearly defined, and the distal leaf margins are incurved.</p>
|tables=
|references=
}}<!--

--><!--

-->{{#Taxon:
name=Grimmia hamulosa
|authority=Lesquereux
|rank=species
|parent rank=subgenus
|synonyms=Grimmia brevirostris
|basionyms=
|family=Grimmiaceae
|habitat=Dry granitic rock and boulders
|elevation=moderate to high elevations (1500-3500 m)
|distribution=Calif.;Oreg.;Wash.
|reference=None
|publication title=Mem. Calif. Acad. Sci.
|publication year=
|special status=
|source xml=https://bibilujan@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/84153f6d59a0a91d69695978a64cee7560374f8e/V27/V27_333.xml
|subfamily=Grimmiaceae subfam. Grimmioideae
|genus=Grimmia
|subgenus=Grimmia subg. Litoneuron
|species=Grimmia hamulosa
|annulus presence=absent
|awn coloration=hyaline
|awn height or length or size=short
|awn presence=absent
|awn shape=not decurrent
|basal marginal laminal cell architecture=thick-walled
|basal marginal laminal cell shape=quadrate;short-rectangular
|basal segment architecture or pubescence or relief=smooth
|base architecture or fixation=clasping
|base shape=tapering
|capsule architecture or pubescence or relief=smooth
|capsule coloration or pubescence or relief=striate
|capsule position=exserted
|capsule presence=absent
|capsule reflectance=shiny
|capsule shape=oblate
|costa size or width=narrow
|distal laminal cell architecture=2-stratose
|distal laminal cell shape=irregularly rounded;quadrate
|distal segment relief=papillose
|intermarginal band presence=absent
|intermarginal band shape=muticous
|laminal cell architecture=thick-walled;sinuose
|laminal cell course=straight
|laminal cell shape=short-rectangular
|leaf length or size=long
|leaf shape=oblong-lanceolate;homomallous-falcate
|margin orientation=erect
|margin shape=plane
|medial laminal cell architecture=thick-walled;sinuose
|medial laminal cell shape=irregularly rounded;quadrate
|operculum shape=conical
|patch prominence or shape=flat
|perichaetial leaf size=not enlarged
|point shape=uncinate;subulate
|point some measurement=2mm;3.5mm
|seta course=straight
|seta some measurement=2.5mm;3.5mm
|stem some measurement=1cm;1.5cm
|stoma presence=absent
|whole_organism coloration=blackish green
|whole_organism growth form=plant
}}<!--

-->[[Category:Treatment]][[Category:Grimmia subg. Litoneuron]]