Cascadia

A. M. Johnson

Amer. J. Bot. 14: 38, figs. 1, 2. 1927 ,.

Etymology: For the Cascade Mountains of western North America
Treatment appears in FNA Volume 8. Treatment on page 48. Mentioned on page 43, 44, 49, 50, 133.

Herbs, annual; herbage glabrous or sparsely red stipitate-glandular. Flowering-stems trailing or suberect, leafy, 4–40 cm. Leaves cauline; stipules absent; petiole glabrous; blade ovate to lanceolate, unlobed, base cuneate or attenuate to rounded, margins 3-toothed apically or not, ultimate margins entire, (without lime-secreting hydathodes), apex acute; venation palmate or pinnate. Inflorescences poorly defined, usually paniculiform or racemiform thyrses, sometimes solitary flowers, 2–4-flowered, bracteate. Flowers: hypanthium adnate to proximal 1/3 of ovary, free from ovary 0 mm, adnate portion increasing along with attached ovary at maturity to constitute 2/3–4/5 of fruit, green, (0.8–2 mm); sepals 5, green; petals 5, white; nectary disc present; stamens 10, (distinct); filaments flattened, slightly widened at base; ovary semi-inferior, 2-locular, carpels connate in proximal 1/3 or less; placentation axile, appearing marginal above point of connation of ovaries; styles 2; stigmas 2. Capsules folliclelike, 2-beaked. Seeds brown, oblong, spiny in longitudinal rows. x = 8.

Distribution

w United States

Discussion

Species 1: w United States.

Johnson placed Saxifraga nuttallii in his monotypic genus Cascadia based on the unusual habit, free carpels, and spiny seeds. Molecular phylogenetic data (M. E. Mort and D. E. Soltis 1999; Soltis et al. 2001) placed Cascadia as sister to the southern South American (Tierra del Fuego) Saxifragodes D. M. Moore, both sister to Micranthes. Mort and Soltis considered the ovary of Cascadia to be superior because the two carpels are distinct to their bases; the hypanthium, fused to each carpel, gives the ovaries a semi-inferior appearance. Ovules in Cascadia are bitegmic, as in Saxifraga; those of Micranthes are usually unitegmic.

Species 1

Selected References

None.

Lower Taxa

"widened" is not a number.

... more about "Cascadia"
epigynous +, perigynous +  and hypogynous +
pinnate +  and palmate +
Luc Brouillet +
A. M. Johnson +
attenuate +  and rounded +
dentate +, serrate +  and entire +
glandular-ciliate +  and ciliate +
unlobed +, ovate +  and lanceolate +
2-beaked +  and folliclelike +
3-carpellate +  and 2-carpellate +
ebracteate +, bracteate +  and 2-300(-1000+)-flowered +
w United States +
For the Cascade Mountains of western North America +
flattened +
bracteate +  and 2-4-flowered +
unisexual +  and bisexual +
4 cm40 mm <br />0.04 m <br /> (40 cm400 mm <br />0.4 m <br />) +
2-3(-4)-beaked +  and folliclelike +
multicellular +
stipitate-glandular +, red +  and glabrous +
free +  and adnate +
racemiform +  and paniculiform +
opposite +  and alternate +
3-toothed +
semi-inferior +
0 cm0 mm <br />0 m <br /> (?) +
crassinucellate +, unitegmic +  and bitegmic +
epigynous +, perigynous +  and hypogynous +
distinct +
Unlobed (?) +  and Lobed (?) +
peltate +  and jointed +
unequal +
Amer. J. Bot. +
straight +
muricate +, cellular-rugulose +, tuberculate +, pitted +, papillate +  and wrinkled +
distinct +
unbranched +  and branched +
persistent +
horizontal +  and erect +
connate +  and distinct +
Cascadia +
Saxifragaceae +
not +, stoloniferous +  and rhizomatous +