Cercocarpus ledifolius var. intermontanus

N. H. Holmgren

Brittonia 39: 424, figs. 1A – D. 1987.

Treatment appears in FNA Volume 9. Treatment on page 338. Mentioned on page 336, 337.

Shrubs or trees, 20–70 (–85) dm. Stems: long-shoot internodes 13–22 mm, sparsely sericeous-villous; short-shoots 2–55 × 1.8–3.4 mm. Leaves: petiole (1.5–) 2.5–6 mm; blade lanceolate to lance-elliptic or (when shorter) elliptic-ovate, (10–) 16–30 (–40) × (2.8–) 5–9 (–11) mm, margins revolute (leaving much of abaxial surface exposed), abaxial surface sericeous or closely villous (hairs not obscuring surface), glabrescent, adaxial sericeous or sparsely villous or hairs tightly coiled and/or 1/2 coiled, glabrescent. Flowers 4–10 per short-shoot; hypanthial tubes strongly sericeous or villous; sepals 2–2.5 mm. Achenes 6.5–10.5 × 1.3–1.5 mm; fruit awns 5–7 (–8.5) cm, proximal setae 2.2–3.5 mm.


Phenology: Flowering May–Jun.
Habitat: Rocky dry, often south- to west-facing slopes, canyons, with Artemisia tridentata, Purshia tridentata, Gambel oak, maple, juniper, pinyon, ponderosa pine, lodgepole pine, Douglas fir and white fir, subalpine zones, often forming pure stands
Elevation: (500–)1300–3300 m

Distribution

V9 554-distribution-map.jpg

Ariz., Calif., Colo., Idaho, Nev., Oreg., Utah, Wash., Wyo., Mexico (Baja California)

Discussion

Variety intermontanus is sympatric with var. ledifolius in southern Idaho, southwestern to eastern Oregon, southeastern Washington, and western Wyoming, and with var. intricatus in northeastern Arizona, eastern California, Nevada, and Utah.

Leaves in individual specimens of var. intermontanus vary in size, with long-shoot leaves largest and the first-formed short-shoot leaves of a season more oblong-ovate and often smaller. Heartwood is hard, very dark, and valued for specialty items. Variety intermontanus has long been known as Cercocarpus ledifolius, or var. ledifolius; its type specimen is convarietal with types of var. intercedens.

Selected References

None.

Lower Taxa

None.
villous +, sericeous +, adaxial +  and glabrescent +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.65 cm6.5 mm <br />0.0065 m <br /> (1.05 cm10.5 mm <br />0.0105 m <br />) +
0.3mm +  and 1.5mm +
1.3mm +  and 1.5mm +
glabrescent +, villous +, hirsute +  and sericeous +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
James Henrickson +  and Brian D. Vanden Heuvel +
N. H. Holmgren +
maroon-brown +, silvery +  and dark gray or brownish black +
compound +  and simple +
opposite +  and alternate +
cuneate +
3 cm30 mm <br />0.03 m <br /> (4 cm40 mm <br />0.04 m <br />) +
5 cm50 mm <br />0.05 m <br /> (7.8 cm78 mm <br />0.078 m <br />) +
0.9 cm9 mm <br />0.009 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
1.6 cm16 mm <br />0.016 m <br /> (3 cm30 mm <br />0.03 m <br />) +
lanceolate +  and lance-elliptic or elliptic-ovate +
0.5 cm5 mm <br />0.005 m <br /> (5 cm50 mm <br />0.05 m <br />) +
5mm +  and 9mm +
free +  and distinct +
hemispheric-campanulate +
Ariz. +, Calif. +, Colo. +, Idaho +, Nev. +, Oreg. +, Utah +, Wash. +, Wyo. +  and Mexico (Baja California) +
not +  and aggregated +
(500–)1300–3300 m +
eglandular +  and veined +
hairy +  and glabrous +
0.1 cm1 mm <br />0.001 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.6 cm6 mm <br />0.006 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
4 +  and 10 +
unisexual +  and bisexual +
not +  and aggregated +
7 cm70 mm <br />0.07 m <br /> (8.5 cm85 mm <br />0.085 m <br />) +
5 cm50 mm <br />0.05 m <br /> (7 cm70 mm <br />0.07 m <br />) +
1.1 cm11 mm <br />0.011 m <br /> (1.35 cm13.5 mm <br />0.0135 m <br />) +
cylindro-fusiform +
0.6 cm6 mm <br />0.006 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
chartaceous-coriaceous +
Rocky dry, often south- to west-facing slopes, canyons, with Artemisia tridentata, Purshia tridentata, Gambel oak, maple, juniper, pinyon, ponderosa pine, lodgepole pine, Douglas fir and white fir, subalpine zones, often forming pure stands +
glabrescent +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0.18 cm1.8 mm <br />0.0018 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
0.26 cm2.6 mm <br />0.0026 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
villous +  and sericeous +
0.3 cm3 mm <br />0.003 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
cinnamon brown +, tinged reddish +  and greenish yellow +
1-3(-18)-flowered +
crassinucellate +
drought-deciduous +  and persistent +
reduced +
flexuous +  and straight +
1.3 cm13 mm <br />0.013 m <br /> (2.2 cm22 mm <br />0.022 m <br />) +
entire +, crenate +, dentate +, flat +  and revolute +
inferior +  and superior +
clustered +, biseriate +  and superposed +
0 (?) +  and 4 (?) +
free +  and distinct +
5 +  and 4 +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
Flowering May–Jun. +
adnate +, free +, connate +  and distinct +
0.22 cm2.2 mm <br />0.0022 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
not arillate +
deciduous +
free +  and distinct +
erect +  and reflexed +
acute-obtuse +  and deltate +
0.2 cm2 mm <br />0.002 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
not +  and branched +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (5.5 cm55 mm <br />0.055 m <br />) +
0.18 cm1.8 mm <br />0.0018 m <br /> (0.34 cm3.4 mm <br />0.0034 m <br />) +
free +  and distinct +
1 +  and 20 +
adnate +  and free +
0.1 cm1 mm <br />0.001 m <br /> (0.38 cm3.8 mm <br />0.0038 m <br />) +
persistent +
distinct +
basal +, lateral +, subterminal +  and terminal +
elongate +
Cercocarpus ledifolius var. intermontanus +
Cercocarpus ledifolius +
variety +
inconspicuous +
hemispheric +
tree +  and shrub +