Cercocarpus ledifolius var. ledifolius

Endemic
Synonyms: Cercocarpus hypoleucus Rydberg C. ledifolius var. intercedens C. K. Schneider
Treatment appears in FNA Volume 9. Treatment on page 337. Mentioned on page 338.

Shrubs or trees, 10–30 dm. Stems: long-shoot internodes 10–17 mm, sparsely villous, sericeous, or hirsute; short-shoots 1–15 × 1.7–3 mm. Leaves: petiole (1.5–) 2–4.5 mm; blade narrowly lanceolate to linear, sometimes lance-elliptic, (7–) 13–29 (–48) × (1.5–) 2–5 (–7) mm, margins revolute (leaving much of abaxial surface exposed), abaxial surface usually densely villous, sometimes sericeous, adaxial usually moderately, often loosely, villous or sericeous, rarely pilose, usually glabrescent. Flowers 1–5 per short-shoot; hypanthial tubes strongly villous; sepals 2–2.5 mm. Achenes 6.5–9 × 1.2–1.4 mm; fruit awns 4–7.5 cm, proximal setae 1.6–2.7 mm.


Phenology: Flowering Apr–May.
Habitat: Rocky outcrops, talus slopes, with Artemisia, Purshia, Gambel oak, pinyon pine and juniper
Elevation: (200–)1300–2300(–3000) m

Distribution

V9 553-distribution-map.jpg

Ariz., Calif., Colo., Idaho, Mont., Nev., Oreg., Utah, Wash., Wyo.

Discussion

Variety ledifolius has narrowly lanceolate to linear leaves that are initially strongly villous adaxially and often strongly villous-woolly abaxially, obscuring the veins between the revolute margins. The adaxial surfaces are usually glabrescent; vestiture is not very dense in sericeous plants.

Here, var. ledifolius comprises intermediates between var. intermontanus and var. intricatus representing hybrids and introgressants that together form breeding populations in zones of parental overlap and in regions outside the range of either parent. Variety ledifolius is considered to be polyphyletic and is much more variable than either parent. Some specimens have relatively small leaves approaching those in var. intricatus. Some specimens have larger long-shoot leaves as in var. intermontanus (with diminished [glabrescent] vestiture and narrower margins) and short-shoot leaves as in var. ledifolius.

Selected References

None.

Lower Taxa

None.
sericeous +  and villous +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.65 cm6.5 mm <br />0.0065 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
0.3mm +  and 1.5mm +
1.2mm +  and 1.4mm +
glabrescent +, pilose +, sericeous +  and villous +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
James Henrickson +  and Brian D. Vanden Heuvel +
Nuttall in J. Torrey and A. Gray +
maroon-brown +, silvery +  and dark gray or brownish black +
compound +  and simple +
opposite +  and alternate +
cuneate +
2.9 cm29 mm <br />0.029 m <br /> (4.8 cm48 mm <br />0.048 m <br />) +
5 cm50 mm <br />0.05 m <br /> (7.8 cm78 mm <br />0.078 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
1.3 cm13 mm <br />0.013 m <br /> (2.9 cm29 mm <br />0.029 m <br />) +
lanceolate to linear +  and lanceolate +
0.5 cm5 mm <br />0.005 m <br /> (5 cm50 mm <br />0.05 m <br />) +
2mm +  and 5mm +
free +  and distinct +
hemispheric-campanulate +
Ariz. +, Calif. +, Colo. +, Idaho +, Mont. +, Nev. +, Oreg. +, Utah +, Wash. +  and Wyo. +
not +  and aggregated +
(200–)1300–2300(–3000) m +
eglandular +  and veined +
hairy +  and glabrous +
0.1 cm1 mm <br />0.001 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.6 cm6 mm <br />0.006 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
1 +  and 5 +
unisexual +  and bisexual +
not +  and aggregated +
4 cm40 mm <br />0.04 m <br /> (7.5 cm75 mm <br />0.075 m <br />) +
1.1 cm11 mm <br />0.011 m <br /> (1.35 cm13.5 mm <br />0.0135 m <br />) +
cylindro-fusiform +
0.6 cm6 mm <br />0.006 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
chartaceous-coriaceous +
Rocky outcrops, talus slopes, with Artemisia, Purshia, Gambel oak, pinyon pine and juniper +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0.18 cm1.8 mm <br />0.0018 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
0.26 cm2.6 mm <br />0.0026 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
cinnamon brown +, tinged reddish +  and greenish yellow +
1-3(-18)-flowered +
crassinucellate +
drought-deciduous +  and persistent +
reduced +
flexuous +  and straight +
hirsute +, sericeous +  and villous +
1 cm10 mm <br />0.01 m <br /> (1.7 cm17 mm <br />0.017 m <br />) +
entire +, crenate +, dentate +, flat +  and revolute +
inferior +  and superior +
clustered +, biseriate +  and superposed +
0 (?) +  and 4 (?) +
free +  and distinct +
5 +  and 4 +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
Flowering Apr–May. +
adnate +, free +, connate +  and distinct +
0.16 cm1.6 mm <br />0.0016 m <br /> (0.27 cm2.7 mm <br />0.0027 m <br />) +
in J. Torrey and A. Gray, Fl. N. Amer. +
not arillate +
deciduous +
free +  and distinct +
erect +  and reflexed +
acute-obtuse +  and deltate +
0.2 cm2 mm <br />0.002 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
not +  and branched +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
0.17 cm1.7 mm <br />0.0017 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
free +  and distinct +
1 +  and 20 +
adnate +  and free +
0.1 cm1 mm <br />0.001 m <br /> (0.38 cm3.8 mm <br />0.0038 m <br />) +
persistent +
distinct +
basal +, lateral +, subterminal +  and terminal +
elongate +
Cercocarpus hypoleucus +  and C. ledifolius var. intercedens +
Cercocarpus ledifolius var. ledifolius +
Cercocarpus ledifolius +
variety +
inconspicuous +
hemispheric +
tree +  and shrub +