Chrysosplenium tetrandrum

Th. Fries

Bot. Not. 1858: 193. 1859 ,.

Common names: Northern golden-saxifrage dorine à quatre étamines
Illustrated
Synonyms: Chrysosplenium alternifolium subsp. tetrandrum (Th. Fries) Hultén
Treatment appears in FNA Volume 8. Treatment on page 74. Mentioned on page 71, 72, 75.

Stolons white, 0.3–1 mm diam., sparsely villous, hairs white, reddish-brown, or purplish. Flowering-stems erect, branching in distal 1/6–1/3 (–1/2), (1.2–) 3–21 cm, glabrous or sparsely villous, especially proximally, hairs white or reddish-brown. Leaves alternate, membranous or fleshy. Stolon leaves: petiole 12–55 mm, usually sparsely villous, sometimes glabrous, hairs white or reddish-brown; blade usually depressed-ovate to reniform, sometimes nearly round, 3–14 × 5–18 mm, base cordate, margins 5–7-crenate, not purple-spotted, glabrous or sparsely ciliate, hairs white, surfaces glabrous or sparsely villous adaxially, hairs usually white, rarely purple. Cauline leaves 1–3 (–4); petiole (2–) 4–20 (–31) mm, glabrous or sparsely villous proximally, hairs white or reddish-brown, sometimes purplish; blade depressed-ovate, flabellate, or reniform, 3–9 × 4–13 mm, base truncate to cuneate or cordate, margins 3–7-crenate, glabrous, surfaces glabrous. Inflorescences terminal, (2–) 3–15-flowered, compact cymes; bracts green, usually purple-spotted, foliaceous, ovate to depressed-ovate or flabellate, 2–9 × 3–12 mm, margins subentire or 3–5-crenate. Pedicels absent or 0.1–1.5 mm. Flowers: hypanthium green or greenish yellow, usually not purple-spotted, rarely purple-spotted distally, turbinate to campanulate, 1–2.2 × 1.2–2.1 mm, glabrous; sepals usually erect, sometimes spreading, usually green, rarely yellow, usually purple-spotted throughout or distally, rarely not purple-spotted, ovate to broadly ovate or depressed-ovate, 0.8–1.5 × 0.8–1.7 mm, apex obtuse to rounded; nectary disc apparently absent; stamens (3–) 4, 0.3–0.4 mm; anthers yellow, 0.1–0.3 × 0.1–0.3 mm; styles 0.2–0.3 mm. Seeds (6–) 15–40, reddish-brown, ovoid to ellipsoid, 0.5–0.8 mm, glabrous. 2n = 24.


Phenology: Flowering May–Aug.
Habitat: Marshy ground, muddy stream banks and shorelines, mossy seeps, bogs, fens
Elevation: 0-3300 m

Distribution

V8 144-distribution-map.gif

Greenland, Alta., B.C., Man., Nfld. and Labr. (Labr.), N.W.T., Nunavut, Ont., Que., Sask., Yukon, Alaska, Colo., Idaho, Mont., Wash., n Eurasia

Discussion

As treated here, Chrysosplenium tetrandrum is circumpolar. In North America, disjunct populations occur in subalpine and alpine habitats in the Bitterroot Range of Idaho and Montana, and in the Front Range of Colorado. Some specimens from northern Europe and the Russian Far East that have been referred to C. alternifolium appear to be morphologically indistinguishable from C. tetrandrum.

Selected References

None.

Lower Taxa

None.

"full" is not a number.

... more about "Chrysosplenium tetrandrum"
epigynous +, perigynous +  and hypogynous +
0.01 cm0.1 mm <br />1.0e-4 m <br /> (0.03 cm0.3 mm <br />3.0e-4 m <br />) +
0.01 cm0.1 mm <br />1.0e-4 m <br /> (0.03 cm0.3 mm <br />3.0e-4 m <br />) +
obtuse +  and rounded +
Craig C. Freeman +  and Nicholas D. Levsen +
Th. Fries +
truncate +  and cuneate or cordate +
depressed-elliptic +
0.3 cm3 mm <br />0.003 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
dentate +, serrate +  and entire +
glandular-ciliate +  and ciliate +
reniform +, flabellate +, depressed-ovate +, round +  and usually depressed-ovate +
0.4 cm4 mm <br />0.004 m <br /> (?) +  and 1.3 cm13 mm <br />0.013 m <br /> (?) +
purple-spotted +  and green +
0.2 cm2 mm <br />0.002 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
ovate;depressed-ovate or flabellate +
0.3 cm3 mm <br />0.003 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
3-carpellate +  and 2-carpellate +
Northern golden-saxifrage +  and dorine à quatre étamines +
ebracteate +, bracteate +  and 2-300(-1000+)-flowered +
Greenland +, Alta. +, B.C. +, Man. +, Nfld. and Labr. (Labr.) +, N.W.T. +, Nunavut +, Ont. +, Que. +, Sask. +, Yukon +, Alaska +, Colo. +, Idaho +, Mont. +, Wash. +  and n Eurasia +
0-3300 m +
lanceolate +  and narrowly oblong +
unisexual +  and bisexual +
1.2 cm12 mm <br />0.012 m <br /> (2 cm20 mm <br />0.02 m <br />) +
2 cm20 mm <br />0.02 m <br /> (30 cm300 mm <br />0.3 m <br />) +
2-3(-4)-beaked +  and folliclelike +
Marshy ground, muddy stream banks and shorelines, mossy seeps, bogs, fens +
multicellular +
purplish +, reddish-brown +  and white +
purple-spotted +, greenish yellow +  and green +
free +  and adnate +
turbinate +  and campanulate +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
1.2mm;2.1mm +
(2-)3-15-flowered +
fleshy +  and membranous +
not purple-spotted +
glabrous +  and ciliate +
3-5-crenate +, subentire +, 3-7-crenate +  and 5-7-crenate +
marginal +, superior +  and inferior +
crassinucellate +, unitegmic +  and bitegmic +
epigynous +, perigynous +  and hypogynous +
distinct +
unlobed +  and lobed +
peltate +  and jointed +
2 cm20 mm <br />0.02 m <br /> (3.1 cm31 mm <br />0.031 m <br />) +
villous +  and glabrous +
0.4 cm4 mm <br />0.004 m <br /> (2 cm20 mm <br />0.02 m <br />) +
Flowering May–Aug. +
unequal +
spiny +  and ribbed +
reddish-brown +
straight +
muricate +, cellular-rugulose +, tuberculate +, pitted +, papillate +  and wrinkled +
ovoid +  and ellipsoid +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
purple-spotted +, yellow +  and green +
distinct +
spreading +  and erect +
ovate +  and broadly ovate or depressed-ovate +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.17 cm1.7 mm <br />0.0017 m <br />) +
Illustrated +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.04 cm0.4 mm <br />4.0e-4 m <br />) +
unbranched +  and branched +
persistent +
horizontal +  and erect +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
connate +  and distinct +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.03 cm0.3 mm <br />3.0e-4 m <br />) +
glabrous +  and villous +
lobed +, truncate +, rounded +  and obtuse +
Chrysosplenium alternifolium subsp. tetrandrum +
Chrysosplenium tetrandrum +
Chrysosplenium +
species +
crenate-dentate +, crenulate +, crenate +  and subentire +
stoloniferous +  and rhizomatous +
annual +, biennial +  and perennial +