Codriophorus sect. Fascicularia

(Bednarek-Ochyra) Bednarek-Ochyra & Ochyra in R. Ochyra et al.

in R. Ochyra et al., Cens. Cat. Polish Mosses, 140. 2003.,.

Basionym: Racomitrium sect. Fascicularia Bednarek-Ochyra Fragm. Florist. Geobot., Ser. Polon. 2: 130. 1995
Treatment appears in FNA Volume 27. Treatment on page 302. Mentioned on page 296, 303.

Stems often subpinnately branched with numerous short, tuft-like, lateral, horizontal branchlets, giving the plants a nodose appearance. Leaves ovatelanceolate, narrowly lanceolate to linear-lanceolate, from an ovate, oblong or ovatelanceolate, not or distinctly plicate base, gradually short or long-acuminate, often with a long, subulate, canaliculate-concave, straight or sometimes wavy to serpentine acumen; margins 1-stratose, recurved to revolute on both sides; apices muticous, acute, subacute to narrowly rounded-obtuse, entire, erose-dentate or denticulate-cristate, epilose or terminated with a hyaline, denticulate awn; costa narrow, vanishing in mid leaf to subpercurrent or percurrent, in the proximal part lying at the bottom of a deep, wide or narrow-angled furrow, open or partly enclosed by the strongly infolded leaf base, in transverse-section 2-stratose throughout, with occasional 3-stratose spots near the base or 2-stratose in the distal and 3–4-stratose in the proximal parts, flattened or convex on the ventral side, not prominently convex, lunate or flattened on the abaxial side, with cells in the abaxial and adaxial rows of similar size and shape or with 1–2 abaxial rows composed of small stereid cells in the proximal 1/2 or at the extreme base; laminal cells 1-stratose, short or long-rectangular to linear, thick and sinuose-walled throughout; basal marginal border differentiated, pellucid; distal laminal cells exceptionally variously 2-stratose and isodiametric.


North America, South America (Argentina), South America (Chile), Europe, arctic and temperate Asia, Atlantic Islands (Azores), Atlantic Islands (Gough Island), Atlantic Islands (Iceland), Atlantic Islands (South Georgia), Atlantic Islands (Tristan da Cunha), Pacific Islands (Hawaiian Islands), Pacific Islands (Society Islands)


Species 6 (3 in the flora).

Members of sect. Fascicularia are recognized by their narrowly lanceolate to linear- or ovate-lanceolate leaves, short-rectangular to linear laminal cells, and peculiar anatomical structure of the costa, which is narrow, usually less than 80 µm wide at the base, and mostly 2-stratose throughout and composed of undifferentiated cells in both adaxial and abaxial rows. Only in the highly isolated C. varius is the costa broader, to 110 µm near the base, and 3–4-stratose in the proximal half, but in all species of the section it is lying at the base of a deep and narrow-angled groove that is often partly enclosed by the strongly folded leaf base.

Selected References



1 Costa 3(-4)-stratose, lunate and distinctly convex on the abaxial side, 75-135 µm wide proximally; leaves usually piliferous, if muticous then entire; peristome teeth 1.0-1.8 mm. Codriophorus varius
1 Costa 2-stratose throughout except the extreme base, flattened and weakly convex on the abaxial side, less than 70 µm wide proximally; leaves muticous; peristome teeth less than 1 mm > 2
2 Costa extending 3/4 way up the leaf to subpercurrent; leaf apex entire; leaf acumen straight Codriophorus fascicularis
2 Costa ending in mid leaf or only slightly distally; leaf apex erose-dentate; leaf acumen serpentine, strongly wavy. Codriophorus corrugatus
wavy +  and serpentine +
Halina Bednarek-Ochyra +  and Ryszard Ochyra +
(Bednarek-Ochyra) Bednarek-Ochyra & Ochyra in R. Ochyra et al. +
both +  and papillose +
denticulate +
short to long +
nodulose;oblate;quadrate;elongate +
thick-walled +  and nodulose-porose +
rectangular;linear +
ovatelanceolate +, oblong +  and ovate +
Racomitrium sect. Fascicularia +
papillose +
undifferentiated +
naked +, plicate +  and smooth +
0 +  and 1/2 +
distinctly verrucose +  and papillose +
conic-mitrate +
small;large +
sulcate +  and smooth +
nearly +  and symmetric +
yellowbrown +, redbrown +  and brown +
striate +  and smooth +
ovoid;long-cylindric +
3-4-stratose;partly enclosed by the strongly infolded leaf base;open;percurrent +
spurred +  and entire +
subpercurrent +
papillose +  and smooth +
flattened +, lunate +  and convex +
much larger +
North America +, South America (Argentina) +, South America (Chile) +, Europe +, arctic and temperate Asia +, Atlantic Islands (Azores) +, Atlantic Islands (Gough Island) +, Atlantic Islands (Iceland) +, Atlantic Islands (South Georgia) +, Atlantic Islands (Tristan da Cunha) +, Pacific Islands (Hawaiian Islands) +  and Pacific Islands (Society Islands) +
sinuose-nodulose +
isodiametric +  and elongate +
narrow-angled +
deep +  and shallow +
wide-angled +
elliptical +  and lingulate +
sinuose-walled +  and 1-stratose +
long-rectangular to linear +  and short +
elliptical +  and oblong-elliptical +
thick-walled +  and sinuose +
dioicous +  and sexual +
curved +  and straight +
long-acuminate +, narrowly lanceolate +  and linear-lanceolate +
erose-dentate to papillose-crenulate or crenate +  and entire +
cristate +  and serrate +
subpercurrent +
long-rostrate +
lanceolate;linear +
in R. Ochyra et al., Cens. Cat. Polish Mosses, +
straight +
erect +, sinistrorse +  and dextrorse +
one;several +
papillose +  and smooth +
short to long +
ornamented +  and smooth +
spheric +
occasional +
creeping +  and ascending +
Racomitrium subg. Cataractarum +
Codriophorus sect. Fascicularia +
Codriophorus +
section +
nodulose +
incrassate +
usually olivaceous;blackish green +
robust +  and gracile +