Codriophorus

P. Beauvois

Mém. Soc. Linn. Paris 1: 445. 1822 ,.

Etymology: Distorted Greek kodon, bell, and phoras, bearing, alluding to capsules with bell-shaped calyptrae
Synonyms: Racomitrium subg. Cataractarum Vilhelm
Treatment appears in FNA Volume 27. Treatment on page 295. Mentioned on page 204, 267, 293, 294.

Plants small, medium to large, mostly stiff and rigid, rarely pliant, robust to gracile, in green, yellow, olive, gray-green, brown to blackish mats. Stems creeping to ascending, repeatedly forked to sparsely fasciculately or pinnately branched, sometimes with short, tuft-like horizontal branchlets. Leaves lanceolate, ovate or oblong-lanceolate, broadly ovate to broadly lingulate, elliptical to oblong-elliptical; margins 1-stratose to variously 2-stratose distally, mostly recurved on one or both sides, entire or erose-dentate to papillose-crenulate or crenate at apex; apices slenderly long-acuminate to rounded, rounded-obtuse to subacute, muticous or seldom with short to long awns, deeply concave to canaliculate-concave; costa single, reaching mid leaf to percurrent, entire or spurred and forked at tip, reniform to strongly flattened abaxially, 2–5-stratose in transverse-section, situated in a shallow or deep, narrow or wide-angled groove; laminal cells 1-stratose throughout to variously 2-stratose distally, elongate to linear throughout or isodiametric to shortly rectangular distally, with sinuose longitudinal walls becoming nodulose and porose with strongly incrassate walls in 2–3 rows at the base, mostly distinctly papillose with large, flat papillae covering longitudinal walls and major parts of the lumina leaving only a narrow groove in the center, occasionally more or less smooth. Perichaetial leaves differentiated, innermost elliptical to lingulate, hyaline throughout or chlorophyllose in the distal part. Seta dextrorse throughout or only with a single torsion to the right proximal to the urn and sinistrorse basally, erect, straight, smooth. Capsule brown, redbrown, or yellowbrown, straight, symmetric or nearly so, ovoid to long-cylindric; exothecial cells isodiametric to elongate, thick-walled; annulus 2–4-seriate; peristome teeth split to base or only to the middle into 2–3 terete, papillose branches. Calyptra distinctly verrucose to papillose distally. Spores spheric, finely papillose.

Distribution

North America, South America (Argentina), South America (Chile), South America (Colombia), Europe, arctic and temperate Asia, n Africa, Atlantic Islands (Azores), Atlantic Islands (Canary Islands), Atlantic Islands (Gough Island), Atlantic Islands (Iceland), Atlantic Islands (Madeira), Atlantic Islands (South Georgia), Atlantic Islands (Tristan da Cunha), Pacific Islands (Hawaiian Islands), Pacific Islands (Society Islands)

Discussion

Species 15 (9 in the flora).

Key

1 Distal laminal cells mostly short and isodiametric; costa robust and broad, usually more than 80 µm wide basally, 2-3-stratose distally and strongly convex abaxially, 3-6-layered in the basal part, lying at the bottom of a shallow, wide-angled groove; leaves lingulate to broadly ovate-lanceolate or lanceolate; plants absent short, lateral tuft-like branchlets. Codriophorus sect. Codriophorus
1 Laminal cells short-rectangular to linear throughout; costa fairly weak and narrow, usually less than 70 µm wide basally, 2-stratose and strongly flattened and weakly convex abaxially throughout or 3(-4)-layered and convex on the abaxial side in the proximal portion of the leaf (C. varius), situated at the bottom of a deep, narrow-angled furrow; leaves narrowly lanceolate to lanceolate; plants often with numerous lateral horizontal tuft-like branchlets Codriophorus sect. Fascicularia
... more about "Codriophorus"
Halina Bednarek-Ochyra +  and Ryszard Ochyra +
P. Beauvois +
both +  and papillose +
toothed +
short to long +
nodulose;oblate;quadrate;elongate +
thick-walled +  and nodulose-porose +
rectangular;linear +
divided +
papillose +
undifferentiated +
naked +, plicate +  and smooth +
0 +  and 1/2 +
distinctly verrucose +  and papillose +
conic-mitrate +
small;large +
sulcate +  and smooth +
nearly +  and symmetric +
yellowbrown +, redbrown +  and brown +
striate +  and smooth +
ovoid;long-cylindric +
2-5-stratose;percurrent +
spurred +  and entire +
subpercurrent +
papillose +  and smooth +
reniform +  and strongly flattened +
much larger +
North America +, South America (Argentina) +, South America (Chile) +, South America (Colombia) +, Europe +, arctic and temperate Asia +, n Africa +, Atlantic Islands (Azores) +, Atlantic Islands (Canary Islands) +, Atlantic Islands (Gough Island) +, Atlantic Islands (Iceland) +, Atlantic Islands (Madeira) +, Atlantic Islands (South Georgia) +, Atlantic Islands (Tristan da Cunha) +, Pacific Islands (Hawaiian Islands) +  and Pacific Islands (Society Islands) +
sinuose-nodulose +
Distorted Greek kodon, bell, and phoras, bearing, alluding to capsules with bell-shaped calyptrae +
isodiametric +  and elongate +
deep +  and shallow +
wide-angled +
elliptical +  and lingulate +
porose +, 2-stratose +  and 1-stratose +
elliptical +  and oblong-elliptical +
thick-walled +  and sinuose +
dioicous +  and sexual +
curved +  and straight +
oblong-lanceolate broadly ovate +  and broadly lingulate +
1-stratose;variously 2-stratose +
erose-dentate to papillose-crenulate or crenate +  and entire +
cristate +  and serrate +
long-rostrate +
lanceolate;linear +
Mém. Soc. Linn. Paris +
bednarek-ochyra2001a +  and bednarek-ochyra2006a +
straight +
erect +, sinistrorse +  and dextrorse +
one;several +
papillose +  and smooth +
short to long +
ornamented +  and smooth +
spheric +
branched +  and fasciculately +
creeping +  and ascending +
Racomitrium subg. Cataractarum +
Codriophorus +
Grimmiaceae subfam. Racomitrioideae +
nodulose +
incrassate +
usually olivaceous;blackish green +
robust +  and gracile +