Cotoneaster villosulus

(Rehder & E. H. Wilson) Flinck & B. Hylmö

Bot. Not. 115: 383. 1962.

Common names: Taiping cotoneaster
Introduced
Basionyms: Cotoneaster acutifolius var. villosulus Rehder & E. H. Wilson in C. S. Sargent, Pl. Wilson 1: 158. 1912 (as acutifolia var. villosula)
Treatment appears in FNA Volume 9. Treatment on page 464. Mentioned on page 452.

Shrubs or trees, 3–5 m. Stems erect, arching, spreading, coarse; branches distichous or spiraled, brown, initially densely villose-strigose. Leaves deciduous; petiole 2–5 mm, strigose; blade elliptic or ovate, 32–72 (–104) x 18–37 (–46) mm, subcoriaceous to chartaceous, base obtuse or cuneate, margins flat, not revolute, veins 5–8, lightly sunken, apex attenuate to long-acuminate, often acute on short-shoots, abaxial surfaces green, shiny, sparsely pilose-strigose, adaxial dark reddish green, shiny, not glaucous, soon dark green, dullish, bulging between lateral-veins, rugose, pilose-strigose; fall leaves purple and rich dark red. Inflorescences on fertile shoots 25–50 mm with 4 leaves, 3–5 (–7) -flowered, mostly compact. Pedicels 3–7 mm, densely strigose. Flowers 6–8 mm, opening small; hypanthium cupulate, tomentose-strigose; sepals: margins tomentose, borders brown, broad, glabrous, apex acute, obtuse, or apiculate, surfaces strigose; petals erect-incurved, pink or red, base dark red, margins white; stamens 18–20, filaments pale-pink, white distally, anthers white; styles 2 or 3. Pomes purple-black, broadly ellipsoid to broadly obovoid, rarely cylindric, 9–11 × 8–11 mm, slightly shiny, not or slightly glaucous, villous; sepals flat or ascending, margins tomentose, strigose; forming star over open navel; style remnants 4/5 from base. Pyrenes 2 or 3.2n = 68 (Germany).


Phenology: Flowering May (often reflowering Sep); fruiting Aug–Nov.
Habitat: Thickets, open forests
Elevation: 0–50 m

Distribution

V9 781-distribution-map.jpg

Introduced; Wash., Asia (China), also in Europe

Discussion

The difficulties of typification and taxonomy for black-fruited species were discussed by J. Fryer and B. Hylmö (1997). Some collections of Cotoneaster villosulus from eastern Washington need study. They approach the very similar C. hsingshangensis J. Fryer & B. Hylmö, a Chinese species with larger, more wrinkled leaves, more globose pomes, and less densely hairy flowers.

Selected References

None.

Lower Taxa

None.

... more about "Cotoneaster villosulus"
dark red +, dark green +, green +  and dark reddish +
pilose-strigose +  and not glaucous +
dullish +  and shiny +
0.4 cm4 mm <br />0.004 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
Jeanette Fryer +, Bertil Hylmö† +  and Peter F. Zika +
(Rehder & E. H. Wilson) Flinck & B. Hylmö +
compound +  and simple +
opposite +  and alternate +
cuneate +  and obtuse +
Cotoneaster acutifolius var. villosulus +
7.2 cm72 mm <br />0.072 m <br /> (10.4 cm104 mm <br />0.104 m <br />) +
15 cm150 mm <br />0.15 m <br /> (21 cm210 mm <br />0.21 m <br />) +
3.7 cm37 mm <br />0.037 m <br /> (4.6 cm46 mm <br />0.046 m <br />) +
3.2 cm32 mm <br />0.032 m <br /> (7.2 cm72 mm <br />0.072 m <br />) +
ovate +  and elliptic +
0.4 cm4 mm <br />0.004 m <br /> (15 cm150 mm <br />0.15 m <br />) +
subcoriaceous +  and chartaceous +
18mm +  and 37mm +
spiraled +  and distichous +
villose-strigose +
adnate +  and distinct +
strigose +  and tomentose +
Taiping cotoneaster +
Wash. +, Asia (China) +  and also in Europe +
not +  and aggregated +
0–50 m +
white +  and pale-pink +
hairy +  and glabrous +
unisexual +  and bisexual +
0.6 cm6 mm <br />0.006 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
not +  and aggregated +
not +  and aggregated +
1.4 cm14 mm <br />0.014 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
orange +  and red or purple +
0.4 cm4 mm <br />0.004 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
Thickets, open forests +
tomentose-strigose +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
flattened +  and domed +
crassinucellate +
deciduous +
pinnate +  and entire +
strigose +  and tomentose +
2 +  and 14 +
inferior +  and superior +
biseriate +  and clustered +
collateral +  and basal +
0.3 cm3 mm <br />0.003 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
0 (?) +  and 4 (?) +
Red (?) +  and Pink (?) +
free +  and distinct +
erect-incurved +
1 +  and 5 +
0.2 cm2 mm <br />0.002 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
Flowering May (often reflowering Sep) +  and fruiting Aug–Nov. +
adnate +, free +, connate +  and distinct +
purple-black +
0.9 cm9 mm <br />0.009 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
villous +  and glaucous +
cylindric +, broadly ellipsoid +  and broadly obovoid +
0.8 cm8 mm <br />0.008 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
trigonous +  and planoconvex +
not arillate +
persistent +
free +  and distinct +
triangular +
2.5 cm25 mm <br />0.025 m <br /> (5 cm50 mm <br />0.05 m <br />) +
tomentose +  and strigose +
Introduced +
free +  and distinct +
branched +  and simple +
spreading +, arching +  and erect +
1 +  and 25 +
short-adnate +
inconspicuous +
lanceolate;linear +
deciduous +
distinct +
lateral +  and terminal +
elongate +
Ostinia +
Cotoneaster villosulus +
Cotoneaster +
species +
inconspicuous +
5 +  and 8 +
tree +  and shrub +