Crataegus pruinosa var. rugosa

(Ashe) Kruschke

Publ. Bot. Milwaukee Public Mus. 3: 181. 1965.

Endemic
Basionym: Crataegus rugosa Ashe J. Elisha Mitchell Sci. Soc. 17: 5. 1900
Synonyms: C. leiophylla Sargent C. mackenziei Sargent ex Mackenzie C. rubicundula Sargent C. seclusa Sargent C. seducta Sargent
Treatment appears in FNA Volume 9. Treatment on page 576. Mentioned on page 575, 577.

Shrubs or trees, 20–40 (–70) dm, usually branched to base. Leaves: blade broadly ovate to deltate, 3–7 cm, length/width = 1–1.2, lobes 2–4 per side, distinct, lobe apex acute, base very broadly cuneate or truncate to weakly subcordate, adaxial surface glabrous, sometimes sparsely appressed-hairy along veins. Inflorescence branches glabrous. Flowers 15–22 mm diam.; stamens 20, anthers pale-pink.


Phenology: Flowering Apr–May; fruiting Sep–Nov.
Habitat: Open scrub, light woodland shade
Elevation: 50–300 m

Distribution

V9 981-distribution-map.jpg

Ont., Ill., Ind., Iowa, Ky., Mich., Mo., N.Y., N.C., Ohio, Pa., Tenn., Wis.

Discussion

Variety rugosa is fairly common from Missouri to the mountains of North Carolina, then northward through the range of the species. The variety often has the most proportionately wide and most deeply incised leaves of the Crataegus pruinosa group. The leaves also characteristically have sharper and longer marginal teeth than other varieties. Extremes are C. mackenziei with acute lobes and C. seducta with rather obtuse lobes.

Unnamed plants of the southern Appalachian piedmont, very likely worthy of at least varietal rank, will key here and are differentiated from var. rugosa as follows: leaf blade 4–6 cm (3–7 cm in var. rugosa), laminas thin (finally chartaceous to somewhat coriaceous in var. rugosa), basal corners of leaf rounded, bases often subcordate (not conspicuously rounded, bases rarely subcordate in var. rugosa).

Selected References

None.

Lower Taxa

None.
appressed-hairy +  and glabrous +
0.8 cm8 mm <br />0.008 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
mauve +  and pink +
James B. Phipps +
(Ashe) Kruschke +
orangebrown +
thin-exfoliating +  and corrugated +
compound +  and simple +
opposite +  and alternate +
truncate +  and weakly subcordate +
Crataegus rugosa +
6 cm60 mm <br />0.06 m <br /> (7 cm70 mm <br />0.07 m <br />) +
mid +  and dark green +
broadly ovate +  and deltate +
3 cm30 mm <br />0.03 m <br /> (7 cm70 mm <br />0.07 m <br />) +
villous +  and glabrous +
adnate +  and distinct +
crowded +, scattered +  and alternate +
Ont. +, Ill. +, Ind. +, Iowa +, Ky. +, Mich. +, Mo. +, N.Y. +, N.C. +, Ohio +, Pa. +, Tenn. +  and Wis. +
not +  and aggregated +
50–300 m +
1.5 cm15 mm <br />0.015 m <br /> (2.2 cm22 mm <br />0.022 m <br />) +
unisexual +  and bisexual +
not +  and aggregated +
not +  and aggregated +
yellow +  and red or purplish +
0.6 cm6 mm <br />0.006 m <br /> (2 cm20 mm <br />0.02 m <br />) +
paler +, gray +, purple-brown +  and reddish-brown +
older +, old +  and new +
Open scrub, light woodland shade +
constricted +
0.2 cm2 mm <br />0.002 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
crassinucellate +
deciduous +
sessile +  and craspedodromous +
glandular-serrate +  and serrate +
subentire +
glabrous +  and short-stipitate-glandular +
subentire +  and entire +
inferior +  and superior +
biseriate +  and clustered +
collateral +  and basal +
0.8 cm8 mm <br />0.008 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
elliptic +  and circular +
0 (?) +  and 4 (?) +
free +  and distinct +
post-mature +
glandular +
Flowering Apr–May +  and fruiting Sep–Nov. +
adnate +, free +, connate +  and distinct +
dark gray-brown +  and russet +
exfoliating +  and corrugated +
pruinose +
scarlet +, bright crimson +  and greenish +
1 cm10 mm <br />0.01 m <br /> (2 cm20 mm <br />0.02 m <br />) +
suborbicular +
Publ. Bot. Milwaukee Public Mus. +
not arillate +
persistent +
free +  and distinct +
triangular +
nonaccrescent +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
1-50-flowered +  and few-leaved +
free +  and distinct +
branched +  and simple +
persistent +
distinct +
exsert +  and lateral +
elongate +
C. leiophylla +, C. mackenziei +, C. rubicundula +, C. seclusa +  and C. seducta +
Crataegus pruinosa var. rugosa +
Crataegus pruinosa +
variety +
6 cm60 mm <br />0.06 m <br /> (10 cm100 mm <br />0.1 m <br />) +
purple +  and shiny black +
straight +
slender +
3 cm30 mm <br />0.03 m <br /> (5 cm50 mm <br />0.05 m <br />) +
craspedodromous +  and gland-tipped +
inconspicuous +
less erect +  and oblique +
1 +  and several +
straight +
determinate +
6 +  and 5 +
tree +  and shrub +