Crataegus pruinosa var. virella

(Ashe) Kruschke

Publ. Bot. Milwaukee Public Mus. 3: 181. 1965.

Endemic
Basionym: Crataegus virella Ashe Ann. Carnegie Mus. 1: 396. 1902
Treatment appears in FNA Volume 9. Treatment on page 577. Mentioned on page 572, 573, 574, 575.

Shrubs or trees, 30–70 dm, larger often single-trunked. Leaves: blade short-ovate to nearly deltate, sometimes ± ovate, 3–4 (–5) cm widest proximal to middle, sometimes near base, length/width = 1.1–1.2, lobes 3 or 4 per side, sinuses shallow, lobe apex acute, base truncate to weakly subcordate (basal corners of leaf rounded), adaxial surface sparsely short-hairy young, usually glabrescent. Inflorescence branches glabrous, sometimes sparsely villous. Flowers 15–22 mm diam.; stamens 20, anthers pale-pink.


Phenology: Flowering Apr–May; fruiting Sep–Nov.
Habitat: Brush, open woodlands
Elevation: 50–300 m

Distribution

V9 982-distribution-map.jpg

Ont., Ark., Mich., Mo., N.Y., Ohio, Pa., Wis.

Discussion

Variety virella is scattered from Arkansas and Missouri to Wisconsin and to Pennsylvania and New York.

The adaxial leaf surface of var. virella is hairy, at least when young. This variety sometimes has hairy pedicels while the leaf blades are among the smallest of the Crataegus pruinosa group; its most commonly found form in recent years, from Missouri to Wisconsin, has proportionately very wide leaves. These are the typical form. The most extreme type possibly belonging to var. virella, the rare C. aspera Sargent [C. mackenziei var. aspera (Sargent) E. J. Palmer] from Missouri, differs by its hairy pedicels, a scabrous hairy adaxial leaf surface as well as permanently densely hairy petioles and veins on abaxial leaf surfaces. It is so densely hairy that it might well pass for C. biltmoreana (ser. Intricatae). However, C. mackenziei var. mackenziei and C. decorata Sargent, usually placed here, possess only scattered hairs on the adaxial surfaces of the young leaves plus some more persistent hairs along the major veins of adaxial surfaces; they are synonymized under var. rugosa and var. pruinosa, respectively.

Selected References

None.

Lower Taxa

None.
glabrescent +  and short-hairy +
0.8 cm8 mm <br />0.008 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
mauve +  and pink +
James B. Phipps +
(Ashe) Kruschke +
orangebrown +
thin-exfoliating +  and corrugated +
compound +  and simple +
opposite +  and alternate +
truncate +  and weakly subcordate +
Crataegus virella +
4 cm40 mm <br />0.04 m <br /> (5 cm50 mm <br />0.05 m <br />) +
mid +  and dark green +
proximal +  and middle +
ovate +, short-ovate +  and nearly deltate +
3 cm30 mm <br />0.03 m <br /> (4 cm40 mm <br />0.04 m <br />) +
villous +  and glabrous +
adnate +  and distinct +
crowded +, scattered +  and alternate +
Ont. +, Ark. +, Mich. +, Mo. +, N.Y. +, Ohio +, Pa. +  and Wis. +
not +  and aggregated +
50–300 m +
1.5 cm15 mm <br />0.015 m <br /> (2.2 cm22 mm <br />0.022 m <br />) +
unisexual +  and bisexual +
not +  and aggregated +
not +  and aggregated +
yellow +  and red or purplish +
0.6 cm6 mm <br />0.006 m <br /> (2 cm20 mm <br />0.02 m <br />) +
paler +, gray +, purple-brown +  and reddish-brown +
older +, old +  and new +
Brush, open woodlands +
constricted +
0.2 cm2 mm <br />0.002 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
villous +  and glabrous +
crassinucellate +
deciduous +
sessile +  and craspedodromous +
glandular-serrate +  and serrate +
subentire +
glabrous +  and short-stipitate-glandular +
subentire +  and entire +
inferior +  and superior +
biseriate +  and clustered +
collateral +  and basal +
0.8 cm8 mm <br />0.008 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
elliptic +  and circular +
0 (?) +  and 4 (?) +
free +  and distinct +
post-mature +
glandular +
Flowering Apr–May +  and fruiting Sep–Nov. +
adnate +, free +, connate +  and distinct +
dark gray-brown +  and russet +
exfoliating +  and corrugated +
pruinose +
scarlet +, bright crimson +  and greenish +
1 cm10 mm <br />0.01 m <br /> (2 cm20 mm <br />0.02 m <br />) +
suborbicular +
Publ. Bot. Milwaukee Public Mus. +
not arillate +
persistent +
free +  and distinct +
triangular +
nonaccrescent +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
1-50-flowered +  and few-leaved +
free +  and distinct +
branched +  and simple +
persistent +
distinct +
exsert +  and lateral +
elongate +
Crataegus pruinosa var. virella +
Crataegus pruinosa +
variety +
6 cm60 mm <br />0.06 m <br /> (10 cm100 mm <br />0.1 m <br />) +
purple +  and shiny black +
straight +
slender +
3 cm30 mm <br />0.03 m <br /> (5 cm50 mm <br />0.05 m <br />) +
craspedodromous +  and gland-tipped +
inconspicuous +
less erect +  and oblique +
1 +  and several +
straight +
determinate +
6 +  and 5 +
tree +  and shrub +