Crataegus sect. Sanguineae

(Zabel ex Rehder) C. K. Schneider

Ill. Handb. Laubholzk. 1: 771. 1906.

Introduced
Basionym: Crataegus unranked Sanguineae Zabel ex Rehder in M. Vilmorin and D. G. J. M. Bois, Frutic. Vilmor., 111. 1904
Treatment appears in FNA Volume 9. Treatment on page 514. Mentioned on page 495, 515, 634.

[Shrubs or] trees, 30–60 [–80] dm. Stems: trunks 1–few, ± erect, bark not recorded; compound thorns on trunks not recorded; thorns on twigs determinate, ± straight and conic, stout. Leaves: blade ± ovate, [3–] 5–9 cm, ± coriaceous, lobes 3–5 per side, sinuses shallow to deep (if deep, usually proximal 1 or 2 lobes only with deep sinuses, with veins to sinuses), veins 4 or 5 [–12] per side, to lobes and deeper sinuses. Inflorescences: branches glabrous [or pubescent]; symmetric bracteoles present, basal bracteoles present, falcate, stipuliform. Flowers: post-mature petals pale paper brown; stamens 20, anthers ivory [pink to purple]. Pomes yellow or orange [red to reddish purple or black]; pyrene sides strongly pitted.

Distribution

Introduced; Wash., c, e Europe, Asia

Discussion

Species ca. 20 (1 in the flora).

Section Sanguineae composes three series ranging continuously from eastern Russia across the whole of southern Siberia and Mongolia to the Pacific, then southward to Afghanistan and the mountains of Pakistan as well as most of eastern China. The section is largely absent from the dry interior tablelands and is disjunct in the Danube basin of central Europe, represented there by Crataegus nigra.

Some of the most diagnostic features of this section are the usually short, stout conic thorns, very dark, one- to two-year twigs, presence of basal falcate bracteoles, long filaments, and laterally erose pyrenes.

Selected References

None.

0.8 cm8 mm <br />0.008 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
James B. Phipps +
(Zabel ex Rehder) C. K. Schneider +
stipuliform +  and falcate +
compound +  and simple +
opposite +  and alternate +
Crataegus unranked +
3 cm30 mm <br />0.03 m <br /> (5 cm50 mm <br />0.05 m <br />) +
5 cm50 mm <br />0.05 m <br /> (9 cm90 mm <br />0.09 m <br />) +
adnate +  and distinct +
crowded +, scattered +  and alternate +
Wash. +, c +, e Europe +  and Asia +
not +  and aggregated +
unisexual +  and bisexual +
not +  and aggregated +
not +  and aggregated +
yellow +  and red or purplish +
0.6 cm6 mm <br />0.006 m <br /> (2 cm20 mm <br />0.02 m <br />) +
tomentose +  and glabrous +
constricted +
0.2 cm2 mm <br />0.002 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
crassinucellate +
deciduous +
entire +  and serrate +
serrate +  and crenate +
inferior +  and superior +
biseriate +  and clustered +
collateral +  and basal +
0.8 cm8 mm <br />0.008 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
elliptic +  and circular +
0 (?) +  and 4 (?) +
free +  and distinct +
post-mature +
adnate +, free +, connate +  and distinct +
dark gray-brown +  and russet +
exfoliating +  and corrugated +
orange +  and yellow +
Ill. Handb. Laubholzk. +
not arillate +
persistent +
free +  and distinct +
appressed +  and erect +
triangular +
1-50-flowered +  and few-leaved +
shallow +  and deep +
Introduced +
free +  and distinct +
branched +  and simple +
persistent +
distinct +
exsert +  and lateral +
elongate +
glabrous +  and tomentose +
Crataegus sect. Sanguineae +
Crataegus +
section +
6 cm60 mm <br />0.06 m <br /> (10 cm100 mm <br />0.1 m <br />) +
straight +
1 cm10 mm <br />0.01 m <br /> (6 cm60 mm <br />0.06 m <br />) +
craspedodromous +  and gland-tipped +
inconspicuous +
1 +  and few +
determinate +
5 +  and 4 +