Cylindropuntia arbuscula

(Engelmann) F. M. Knuth in C. Backeberg and F. M. Knuth

in C. Backeberg and F. M. Knuth, Kaktus-ABC, 123. 1935.

Common names: Pencil cholla
Basionym: Opuntia arbuscula Engelmann Proc. Amer. Acad. Arts 3: 309. 1856
Treatment appears in FNA Volume 4. Treatment on page 106. Mentioned on page 104, 109.

Trees or shrubs, densely branched, 0.5–3 m. Stem segments green, sometimes purple-tinged, 6–10.5 × 0.5–1.3 cm; tubercles narrowly elongate, appearing as wrinkles when dry, 1–2 cm; areoles circular, 2–3 mm in diam.; wool tan to brown, aging gray to black. Spines 0–2 (–3) per areole, sparsely distributed along stem, usually deflexed, pale-yellow or redbrown aging black, stout, the longest 0.8–3.5 (–5) cm; sheaths loose fitting, yellowish-brown. Glochids in adaxial tuft and marginal, encircling areole, pale-yellow. Flowers: inner tepals green or orange-bronze, spatulate, 17–20 mm, apiculate; filaments dark green-bronze; anthers yellow; style whitish basally to light orange apically; stigma lobes very pale green. Fruits green, becoming yellowish apically, sometimes tinged red to purplish at areoles, commonly sterile, then narrow, tuberculate, to 2.5 cm, fertile ones stipitate, obconic, 20–50 × 15–35 mm, fleshy, becoming smooth, spineless; umbilicus 3–4 mm deep; areoles 15–17 (–34). Seeds pale-yellow, angularly circular to oblong, thick, 3–5 mm diam., lumpy; girdle smooth, broad, not protruding. 2n = 66.


Phenology: Flowering spring (Apr–Jun).
Habitat: Sonoran Desert flats, bajadas, desert scrub
Elevation: 300-1000 m

Discussion

Cylindropuntia arbuscula forms hybrids with C. leptocaulis in south-central Arizona; those hybrids have narrow, obscurely tuberculate stems and reddish fruits nearly the size of those of C. arbuscula and chromosome number of 2n = 55. Hybrids between C. arbuscula and C. spinosior [= C. ×neoarbuscula (Griffiths) F. M. Knuth] have large green fruits, which often split open, and on stem segments distal areoles that usually bear three or four spines to 2.5 cm. Hybrids between C. arbuscula and C. versicolor [= C. ×vivipara (Rose) F. M. Knuth] have large green fruits, which often split open, and distal stem segment areoles that bear one or two spines to 1.5 cm.

Selected References

None.

Lower Taxa

None.

"thick" is not a number.

... more about "Cylindropuntia arbuscula"
cushionlike +
gray +  and tan +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
deciduous +  and persistent +
15 +  and 17 +
obdeltate +  and rhombic +
Donald J. Pinkava +
(Engelmann) F. M. Knuth in C. Backeberg and F. M. Knuth +
Opuntia arbuscula +
proliferating +
flattened +  and cylindric +
Pencil cholla +
Ariz. +  and Mexico (Sonora) +
300-1000 m +
dark green-bronze +
15 cm150 mm <br />0.15 m <br /> (30 cm300 mm <br />0.3 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (15 cm150 mm <br />0.15 m <br />) +
spineless +  and stipitate +
tinged red;purplish +
indehiscent +
persistent +, long +  and deciduous +
20mm;50mm +
tuberculate +
fertile +  and sterile +
obconic +
0 cm0 mm <br />0 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
not protruding +
Sonoran Desert flats, bajadas, desert scrub +
1.7 cm17 mm <br />0.017 m <br /> (2 cm20 mm <br />0.02 m <br />) +
deciduous +
flattened +  and conic +
0 cm0 mm <br />0 m <br /> (3 cm30 mm <br />0.03 m <br />) +
succulent +
tuberculate +
fluted;spheric;depressed-spheric or club-shaped +
tinged color +
sometimes covered by outgrowths of proximal portion of style base or of flower tube wall +  and open +
persistent +  and deciduous +
Flowering spring (Apr–Jun). +
ridgelike +  and nipple--shaped +
in C. Backeberg and F. M. Knuth, Kaktus-ABC, +
3;2;2;30 +
tuberlike +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
angularly circular;oblong +
fitting +
distributed +
3.5 cm35 mm <br />0.035 m <br /> (5 cm50 mm <br />0.05 m <br />) +
black +, redbrown +  and pale-yellow +
deciduous +  and persistent +
conspicuous +
0 (?) +  and 2 (?) +
flattened +
0.8 cm8 mm <br />0.008 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
hard +  and rigid +
decurrent +
winged +  and unsegmented +
tuberculate +  and smooth +
40 cm400 mm <br />0.4 m <br /> (50 cm500 mm <br />0.5 m <br />) +
purple-tinged +  and green +
6 cm60 mm <br />0.06 m <br /> (10.5 cm105 mm <br />0.105 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (1.3 cm13 mm <br />0.013 m <br />) +
segmented +
woody +  and succulent +
light orange +  and whitish +
Opuntia sect. Cylindropuntia +
adventitious +
tuberlike +
epipetric +  and epiphytic +
Cylindropuntia arbuscula +
Cylindropuntia +
species +
dull +  and glossy +
0.3 cm3 mm <br />0.003 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
shrub +  and tree +
terrestrial;epiphytic or epipetric +
gray +  and black +
1 cm10 mm <br />0.01 m <br /> (2 cm20 mm <br />0.02 m <br />) +