Cyrtomium

C. Presl

Tent. Pterid. 86. 1836.

Common names: Asiatic holly fern
Etymology: Greek cyrtoma, arch, for the arched veins
Treatment appears in FNA Volume 2.

Plants terrestrial or on rock. Stems erect or ascending, stolons absent. Leaves monomorphic, evergreen. Petiole ± 1/2–3/4 length of blade, base not swollen; vascular-bundles more than 3, arranged in arc, ± round in cross-section. Blade oblong-lanceolate, 1-pinnate with pinnae not lobed, distal pinnae only slightly smaller, blade ending in basally lobed pinna ± similar to lateral pinnae, papery or somewhat leathery. Pinnae not articulate to rachis, segment margins crenate to spinulose, sometimes also coarsely dentate; proximal pinnae not reduced, petiolulate, equilateral or somewhat inequilateral with acroscopic base more developed; costae adaxially grooved, grooves continuous from rachis to costae; indument of filiform scales abaxially, absent adaxially. Veins elaborately anastomosing, areoles formed with 1–3 included veinlets. Sori in 2 or more rows between midrib and margin, round. Indusia peltate, persistent or caducous. Spores brown, with inflated folds or wings. x = 41.

Distribution

North America, Asia, Africa including Madagascar, Pacific Islands (Hawaii)

Discussion

This difficult genus requires further systematic study. Estimates of species numbers have ranged from 9 (C. Christensen 1930) to 59 (Shing K. H. 1965). The group might better be considered a subgenus of Polystichum, from which it is poorly differentiated morphologically.

Species ca. 15 (2 in the flora).

Key

1 Pinnae 4-10(-12) pairs, leathery, shiny adaxially, margins sometimes undulate or coarsely dentate, but not minutely crenulate or denticulate. Cyrtomium falcatum
1 Pinnae (8-)10-25 pairs, papery, not shiny adaxially, margins minutely crenulate-denticulate. Cyrtomium fortunei var. fortunei
... more about "Cyrtomium"
George Yatskievych +
C. Presl +
not swollen +
simple to commonly +
oblong-lanceolate +
roundish +
Asiatic holly fern +
North America +, Asia +, Africa including Madagascar +  and Pacific Islands (Hawaii) +
Greek cyrtoma, arch, for the arched veins +
inflated +
often +  and glabrous +
caducous +  and persistent +
monomorphic +
evergreen +
1/2 length of blade;3/4 length of blade +
not articulate +
Lobed (?) +  and Not lobed (?) +
inequilateral +
Tent. Pterid. +
round +, cuplike +, hoodlike +, reniform +, falcate +  and linear +
ching1936a +, christensen1930a +  and shing1965a +
2 (?) +  and 3 (?) +
persistent +
filiform +
crenate to spinulose +
discrete +
variously--shaped +
reniform +  and oblong +
dictyostelic +, unbranched +  and branched +
ascending +  and erect +
Cyrtomium +
Dryopteridaceae +
anastomosing +
on rock +  and terrestrial +
epiphytic +  and hemiepiphytic +