Dichanthelium commutatum

(Schult.) Gould
Common names: Variable panicgrass
Synonyms: Panicum commutatum Panicum divergens
Treatment appears in FNA Volume 25. Treatment on page 414.

Plants cespitose, with caudices or with rhizomes up to 2 mm thick. Basal rosettes well-differentiated; blades 1-14 cm long, to 22 mm wide, ovate to lanceolate. Culms 20-75 cm, erect or decumbent to sprawling, often purplish; nodes and internodes glabrous or puberulent to pubescent; fall phase initially nearly erect, often sprawling eventually, branches initially erect and apparently dichotomous, later rebranching, blades and secondary panicles smaller than those of the culms. Cauline leaves 4-6; sheaths not overlapping, often glaucous, purplish, or olivaceous, glabrous or puberulent, margins usually ciliate; ligules about 0.3 mm, membranous, ciliate, cilia longer than the membranous portion, rarely with adjacent, about 12 mm hairs; blades 5-16 cm long, 5-25 mm wide, linear to ovatelanceolate, glabrous or puberulent, with 9-13 major veins and 30-80 minor veins, bases cordate-clasping, often asymmetrical, with papillose-based marginal cilia. Panicles 5-12 cm long, 3-10 cm wide, open, exserted; branches flexuous. Spikelets 2.2-3.2 mm long, 1.1-1.3 mm wide, ellipsoid, yellowish-green or purplish, pubescent. Lower glumes 0.7-1.8 mm; upper glumes and lower lemmas equaling or slightly shorter than the spikelets; lower florets sterile; upper florets often minutely umbonate. 2n = 18.

Distribution

Del., W.Va., Fla., Conn., D.C., N.J., Mass., R.I., La., Tenn., N.C., S.C., Pa., N.Y., Va., Ala., Ark., Ill., Ga., Ind., Maine, Md., Tex., Ohio, Mo., Mich., Okla., Miss., Ky.

Discussion

Dichanthelium commutatum is fairly common in dry to wet, semi-open woodlands. Its range extends from the eastern United States to South America. The primary panicles are open-pollinated and are produced from April through June; the secondary panicles are primarily cleistogamous and are produced from June through fall. The four subspecies are fairly distinct in some parts of their ranges, but subsp. commutatum intergrades with the other three where they occur together.

Selected References

None.

Key

1 Culms densely crisp-puberulent; spikelets 2.2-2.7 mm long; cauline blades usually 5-8 cm long, 5-10 mm wide, thick, the bases symmetrical; rosette blades usually less than 3 cm long and to 6 mm wide Dichanthelium commutatum subsp. ashei
1 Culms usually glabrous or sparsely pubescent; spikelets 2.6-3.2 mm long; cauline blades usually more than 8 cm long and 10 mm wide, thin, bases sometimes asymmetrical; rosette blades large, some more than 4 cm long and 10 mm wide. > 2
2 Cauline blades nearly linear, 5-14 mm wide, about 10 times as long as wide; spikelets 3-3.2 mm long; lower glumes about 1/2 as long as the spikelets Dichanthelium commutatum subsp. equilaterale
2 Cauline blades ovate-lanceolate, 6-25 mm wide, about 4-8 times as long as wide; spikelets 2.6-3.2 mm long; lower glumes about 1/4 as long as the spikelets. > 3
3 Culms decumbent or sprawling, with loose caudices or rhizomes; blades strongly asymmetric-falcate, often; spikelets 2.9-3.2 mm long; lower lemmas pointed Dichanthelium commutatum subsp. joorii
3 Culms more or less erect, with caudices; blades almost symmetrical, green, sometimes glaucous; spikelets 2.6-2.9 mm long; lower lemmas rounded Dichanthelium commutatum subsp. commutatum

"usually distinctly longer and narrower" is not a number."decumbent" is not a number.

... more about "Dichanthelium commutatum"
rounded +  and attenuate +
anatomy +  and kranz +
Mary E. Barkworth +
(Schult.) Gould +
1 (?) +  and 3 (?) +
keeled +  and rounded +
well-differentiated +
pseudopetiolate +  and branching +
divergent +  and parallel +
intravaginal +, extravaginal +, branching +  and basal +
swelling +
rebranching +  and dichotomous +
non-radiate +  and radiate +
distinct +
5 cm50 mm <br />0.05 m <br /> (16 cm160 mm <br />0.16 m <br />) +
puberulent +  and glabrous +
linear +  and ovatelanceolate +
0.5 cm5 mm <br />0.005 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
spikelike +
0 cm0 mm <br />0 m <br /> (1 cm10 mm <br />0.01 m <br />) +
not developed +
terete +  and compressed +
12 cm120 mm <br />0.12 m <br /> (?) +
Variable panicgrass +
rebranching +
not dense +
decumbent +  and sprawling +
decumbent to sprawling +  and erect +
20 cm200 mm <br />0.2 m <br /> (75 cm750 mm <br />0.75 m <br />) +
Del. +, W.Va. +, Fla. +, Conn. +, D.C. +, N.J. +, Mass. +, R.I. +, La. +, Tenn. +, N.C. +, S.C. +, Pa. +, N.Y. +, Va. +, Ala. +, Ark. +, Ill. +, Ga. +, Ind. +, Maine +, Md. +, Tex. +, Ohio +, Mo. +, Mich. +, Okla. +, Miss. +  and Ky. +
capillary +
germination +
pistillate +  and staminate +
sometimes longer +
compressed +  and terete +
shorter or longer +
subtending +
unequal +
membranous +
papillose-based +
uncinate +
0.1 cm1 mm <br />0.001 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
Oval (?) +  and Round (?) +
bracteate +  and ebracteate +
solid +  and hollow +
puberulent +  and pubescent +
cauline +  and basal +
coriaceous +
0.3 cm3 mm <br />0.003 m <br /> (?) +
membranous +
inconspicuous +
1/5 +  and 3/4 +
acuminate +, acute +  and truncate +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.18 cm1.8 mm <br />0.0018 m <br />) +
complex +  and simple +
concealed +  and prominent +
puberulent +  and pubescent +
2 +  and 1 +
5cm +  and 12cm +
subterranean +
3 cm30 mm <br />0.03 m <br /> (10 cm100 mm <br />0.1 m <br />) +
dry +  and fleshy +
membranous +
rebranching +  and dichotomous +
rebranching +  and dichotomous +
not overlapping +
not viscid +
puberulent +, glabrous +  and glaucous +
in triplets +  and paired +
purplish +  and yellowish-green +
0.22 cm2.2 mm <br />0.0022 m <br /> (0.32 cm3.2 mm <br />0.0032 m <br />) +
1 +  and 60 +
unisexual +  and bisexual +
ellipsoid +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.52 cm5.2 mm <br />0.0052 m <br />) +
1.1mm +  and 1.3mm +
1 +  and 3 +
fused +  and free +
2 +  and 3 +
dome--shaped +  and triangular +
Panicum commutatum +  and Panicum divergens +
Dichanthelium commutatum +
Dichanthelium sect. Macrocarpa +
species +
membranous +
chartaceous-indurate +
30 (?) +  and 80 (?) +
plant +  and cespitose +
aquatic +  and terrestrial +
dioecious +, monoecious +  and synoecious +