Dryas ajanensis subsp. beringensis

Jurtzev in A. I. Tolmatchew

in A. I. Tolmatchew, Fl. Arct. SSSR 9(1): 322. 1984.

Common names: White dryad
Illustrated
Synonyms: Dryas octopetala var. kamtschatica (Juzepczuk) Hultén D. octopetala var. luteola Hultén D. octopetala var. viscida Hultén
Treatment appears in FNA Volume 9. Treatment on page 329. Mentioned on page 327.

Plants 2.5–9 cm. Leaf-blades oblong or ovate to elliptic, 3–10 × 0.5–4 mm, base usually truncate or cordate, sometimes cuneate, margins flat, coarsely dentate to serrate, sinuses 5–25% to midvein, apex acute to rounded, surfaces plicate along midvein and/or rugulose or rugose, midvein and lateral-veins sunken, abaxial tomentose to woolly, adaxial usually glabrous, sometimes sparsely hairy proximally on midvein, feathery hairs abundant along midveins abaxially and on petioles, rarely scattered elsewhere, midveins and petioles abaxially usually stipitate-glandular. Peduncles 38–50 mm. Flowers erect at flowering; sepals linear-oblong, 4–6 × 1.5–2 mm; petals 8, spreading, white or cream, 9–14 × 5–11 mm; filaments glabrous. Achenes 2.5–3 mm; styles 16–38 mm. 2n = 18 (Chukotka, Siberia).


Phenology: Flowering Jun–Aug.
Habitat: Alpine heath, dry slopes, timberline, moraines, tundra
Elevation: 0–1800 m

Distribution

V9 540-distribution-map.jpg

N.W.T., Yukon, Alaska, e Asia (Russian Far East: Chukotka)

Discussion

Subspecies beringensis was formerly considered to be part of Dryas octopetala var. octopetala (R. Elven et al., http://nhm2.uio.no/paf/). This subspecies forms hybrid swarms with D. alaskensis and D. integrifolia in Alaska and eastern Siberia (Elven et al.). Dryas incanescens Juzepczuk and D. ajanensis Juzepczuk var. incanescens (Juzepczuk) Kozhevnikov have been misapplied to subsp. beringensis.

Selected References

None.

Lower Taxa

None.
tomentose +  and woolly +
2.5mm +  and 3mm +
hairy +  and glabrous +
Joshua C. Springer +  and Bruce D. Parfitt† +
Jurtzev in A. I. Tolmatchew +
blackish +, reddish +  and brown +
compound +  and simple +
opposite +  and alternate +
cuneate +, cordate +  and truncate +
3.9 cm39 mm <br />0.039 m <br /> (4.6 cm46 mm <br />0.046 m <br />) +
smooth +  and rugose +
ovate +, oval +, obovate +, oblongelliptic +, oblong +, linear +, lanceolate +  and elliptic +
0.2 cm2 mm <br />0.002 m <br /> (3.9 cm39 mm <br />0.039 m <br />) +
stoloniferous +
free +  and distinct +
White dryad +
N.W.T. +, Yukon +, Alaska +  and e Asia (Russian Far East: Chukotka) +
not +  and aggregated +
0–1800 m +
stipitate-glandular +  and villous +
1.3 cm13 mm <br />0.013 m <br /> (2.9 cm29 mm <br />0.029 m <br />) +
unisexual +  and bisexual +
not +  and aggregated +
linear +  and lanceolate +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
Alpine heath, dry slopes, timberline, moraines, tundra +
0.3 cm3 mm <br />0.003 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
crassinucellate +
0.3 cm3 mm <br />0.003 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
persistent +
crenate +  and 1-2-toothed +
stipitate-glandular +
rugose +  and rugulose +
inferior +  and superior +
clustered +, biseriate +  and superposed +
3.8 cm38 mm <br />0.038 m <br /> (5 cm50 mm <br />0.05 m <br />) +
persistent +  and deciduous +
free +  and distinct +
0.9 cm9 mm <br />0.009 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
stipitate-glandular +
Flowering Jun–Aug. +
adnate +, free +, connate +  and distinct +
in A. I. Tolmatchew, Fl. Arct. SSSR +
not arillate +
persistent +
free +  and distinct +
erect-ascending +, ascending +  and spreading +
linear-oblong +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
Illustrated +
free +  and distinct +
rooting +  and branched +
adnate +  and free +
persistent +
distinct +
basal +, lateral +, subterminal +  and terminal +
elongate +
1.6 cm16 mm <br />0.016 m <br /> (3.8 cm38 mm <br />0.038 m <br />) +
Dryas octopetala var. kamtschatica +, D. octopetala var. luteola +  and D. octopetala var. viscida +
Dryas ajanensis subsp. beringensis +
Dryas ajanensis +
subspecies +
inconspicuous +
hemispheric +