Drymocallis pseudorupestris var. crumiana

D. D. Keck ex Ertter

J. Bot. Res. Inst. Texas 1: 39, figs. 1G–L. 2007.

Common names: Crum’s drymocallis or wood beauty
Endemic
Treatment appears in FNA Volume 9. Treatment on page 290. Mentioned on page 281, 289.

Stems (0.3–) 0.8–2 (–2.5) dm, base 1–2 mm diam., short hairs absent or sparse. Basal leaves (2–) 3–9 (–15) cm, not to sparsely short-hairy, sparsely to moderately subsessile golden glandular, rigidly bristly (bristles 0.5–1 mm); leaflet pairs 3–4 (–5); terminal leaflet broadly obovate to flabellate, 0.2–1.2 (–2) × 0.5–1.5 cm, teeth ± single, 2–5 per side. Pedicels 5–12 (proximal to 20) mm, bristly, short hairs absent or sparse. Flowers 2–8; hypanthia and sepals prominently bristly, bristles 1–1.5 mm; epicalyx bractlets elliptic-ovate to lanceolate, 2–3.5 (–4) × 1–2 mm; sepals 4–6 (–7) mm; petals not or scarcely overlapping, often red-tinged, narrowly to broadly obovate, 5–7 (–9) × 3–5 mm; filaments 1.5–2 (–3) mm; styles usually dark red, rarely golden brown.


Phenology: Flowering Jul–Aug.
Habitat: Rocky slopes, talus, and ledges, on metamorphic, granitic, and volcanic substrates
Elevation: 3200–3900 m

Discussion

The red-tinged petals of var. crumiana, initially noted by E. Crum (as summarized by B. Ertter 2007), are not reliably diagnostic; however, the distinctive leaflet aspect is not found elsewhere in Drymocallis, though approaching D. fissa in Colorado. The vestiture includes prominent golden glands and conspicuous bristles that are rarely found in other North American Drymocallis, and the stigmas are commonly (though not consistently) dark reddish, not the more common pale golden brown. The variety occurs in the southern Sierra Nevada in Fresno, Inyo, and Tulare counties, and in the White Mountains of Mono County.

Selected References

None.

Lower Taxa

None.

"dm" is not declared as a valid unit of measurement for this property.

light-brown +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
Barbara Ertter +
D. D. Keck ex Ertter +
compound +  and simple +
opposite +  and alternate +
9 cm90 mm <br />0.09 m <br /> (15 cm150 mm <br />0.15 m <br />) +
3 cm30 mm <br />0.03 m <br /> (9 cm90 mm <br />0.09 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
Potentilla pseudorupestris +
oblanceolate-oblong +
1 cm10 mm <br />0.01 m <br /> (25 cm250 mm <br />0.25 m <br />) +
reduced +
(10-)20-40(-50)° +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
free +  and distinct +
Crum’s drymocallis or wood beauty +
not +  and aggregated +
3200–3900 m +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
elliptic-ovate +  and lanceolate +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
2 +  and 8 +
unisexual +  and bisexual +
not +  and aggregated +
deciduous +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
Rocky slopes, talus, and ledges, on metamorphic, granitic, and volcanic substrates +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
open +, leafy +  and 2-40-flowered +
1/6 +  and 3/4-4/5 +
crassinucellate +
opposite +  and alternate +
marcescent +
winter-persistent +
hairy +  and glabrate +
overlapping +  and not confluent +
elliptic-ovate +  and flabellate or rhombic +
toothed +, or +, flat +, entire +  and more or less toothed +
inferior +  and superior +
clustered +, biseriate +  and superposed +
0.5 cm5 mm <br />0.005 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
overlapping +
0.7 cm7 mm <br />0.007 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
0 (?) +  and 4 (?) +
free +  and distinct +
0.5 cm5 mm <br />0.005 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
Flowering Jul–Aug. +
adnate +, free +, connate +  and distinct +
J. Bot. Res. Inst. Texas +
not arillate +
0.6 cm6 mm <br />0.006 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
persistent +
free +  and distinct +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
free +  and distinct +
branched +  and simple +
openly tufted +  and loosely spaced +
stramineous +, reddish +  and green +
1 +  and 20 +
palmate +  and pinnate +
golden brown +  and dark red +
deciduous +
distinct +
fusiform +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
Drymocallis glandulosa subsp. pseudorupestris +, P. glandulosa subsp. pseudorupestris +, P. glandulosa var. pseudorupestris +  and P. rupestris var. americana +
Drymocallis pseudorupestris var. crumiana +
Drymocallis pseudorupestris +
variety +
1.2 cm12 mm <br />0.012 m <br /> (2 cm20 mm <br />0.02 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
broadly obovate +  and flabellate +
0.5 cm5 mm <br />0.005 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
horseshoe--shaped +
2 +  and 5 +
inconspicuous +
hemispheric +  and conic +
enlarged +
peglike-glandular +  and septate-glandular +