Elymus diversiglumis

Scribn. & C.R. Ball.
Common names: Unequal-glumed wildrye
Endemic
Treatment appears in FNA Volume 24. Treatment on page 316.

Plants cespitose, not rhizomatous, sometimes moderately glacous. Culms 70-160 cm, erect; nodes 4-9, mostly exposed, glabrous. Leaves evenly distributed; sheaths glabrous, often purplish; auricles 1-2 mm, purplish or brownish black; ligules usually 1-2 mm; blades 5-17 mm wide, lax, adaxial surfaces usually pilose, at least on the veins, occasionally scabrous. Spikes 8-28 cm long, 3-5 cm wide, nodding to pendent, with 2 spikelets per node, rarely with 1 or 3 at a few nodes; internodes 4-6 (9) mm long, 0.2-0.3 mm thick at the thinnest sections, margins and summits often pubescent. Spikelets 10-16 mm, appressed, with 2-4 (5) florets, lowest florets functional; disarticulation above the glumes, beneath each floret. Glumes usually differing in length by at least (3) 4 mm, occasionally obsolete, (1) 2-15 (20) mm long including the undifferentiated awns, indurate at the base, (0.1) 0.2-0.4 (0.6) mm wide, setaceous, 0-1-veined, tapering from the base, scabrous or hispidulous at least towards the apices, margins firm, awns often outcurving; lemmas 7-12 mm, usually silvery-hirsute to sericeous, occasionally hirtellous or strigose, at least near the margins, backs sometimes scabrous, awns 18-35 mm, moderately to strongly outcurving at maturity; paleas 7-10 mm, obtuse, occasionally emarginate; anthers 2-4 mm. Anthesis from early June to late July. 2n = 28.

Distribution

Okla., Man., Ont., Sask., Minn., Mich., N.Dak., S.Dak., Wis., Ark., Ill., Iowa, Mo., Wyo.

Discussion

Elymus diversiglumis grows in moist to dry, often base-rich and alluvial soils, in open woods, woodland margins, and thickets in the northern Great Plains, from Saskatchewan and Manitoba to Wyoming, Wisconsin, and Iowa.

Elymus diversiglumis is a variable species that, like E. svensonii (p. 314) and E. churchii (p. 314), may have originated from hybrids between E. canadensis var. canadensis (p. 305) and E. hystrix (see next), although part of its range extends further west than the current distribution of the latter species. Elymus diversiglumis usually reaches anthesis 2-4 weeks earlier than sympatric populations of E. canadensis. Church (1954, 1958, 1967a) found that most artificial canadensis-hystrix hybrids, as well as some plants of E. diversiglumis itself, are sterile. Those that were not sterile could occasionally form fertile backcrosses with E. canadensis and, to a lesser extent, with E. hystrix. Introgressant populations involving all three species are known. Artificial crosses with other species have not been successful.

Selected References

None.

Lower Taxa

None.

"decumbent" is not a number.

... more about "Elymus diversiglumis"
membranous +  and scarious +
0.2 cm2 mm <br />0.002 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
bidentate +  and acute +
brownish black +  and purplish +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
Mary E. Barkworth +, Julian J.N. Campbell +  and Bjorn Salomon +
Scribn. & C.R. Ball. +
1 (?) +  and 3 (?) +
straight +
outcurving +
basal +  and apical +
from the sinus +  and terminal +
undifferentiated +
not geniculate +
1.8 cm18 mm <br />0.018 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
keeled +  and rounded +
pseudopetiolate +  and branching +
intravaginal +, extravaginal +, branching +  and basal +
swelling +
0.2mm +  and 0.4mm +
not evident +
cross +, linear +  and narrowly lanceolate +
0.5 cm5 mm <br />0.005 m <br /> (1.7 cm17 mm <br />0.017 m <br />) +
spikelike +
0 cm0 mm <br />0 m <br /> (1 cm10 mm <br />0.01 m <br />) +
hairy +  and glabrous +
Unequal-glumed wildrye +
not branching +
sometimes strongly decumbent +  and prostrate +
70 cm700 mm <br />0.7 m <br /> (160 cm1,600 mm <br />1.6 m <br />) +
not woody +
Okla. +, Man. +, Ont. +, Sask. +, Minn. +, Mich. +, N.Dak. +, S.Dak. +, Wis. +, Ark. +, Ill. +, Iowa +, Mo. +  and Wyo. +
liquid +, soft +  and hard +
capillary +
pistillate +  and staminate +
2 +  and 4 +
laterally compressed +  and terete +
shorter than to longer than the adjacent florets +
2mm +  and 15mm +
subtending +
hispidulous +  and scabrous +
tapering +  and setaceous +
unequal +, subequal +  and reduced +
uncinate +
solid +  and hollow +
1.5 cm15 mm <br />0.015 m <br /> (?) +
4mm +  and 6mm +
0.2 cm2 mm <br />0.002 m <br /> (2.6 cm26 mm <br />0.026 m <br />) +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.03 cm0.3 mm <br />3.0e-4 m <br />) +
ciliate +  and scabrous +
distributed +
usually silvery-hirsute +  and sericeous occasionally hirtellous or strigose +
linear-lanceolate +
0.7 cm7 mm <br />0.007 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
stiffly membranous +  and coriaceous +
erose +  and entire +
acute +, rounded +  and truncate +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
membranous +
inconspicuous +
lanceolate +
fleshy +  and membranous +
few +, 1 +, 4 +  and 9 +
2 +  and 1 +
hairy +  and glabrous +
well-developed +
slightly longer +
subequal +
0.7 cm7 mm <br />0.007 m <br /> (1 cm10 mm <br />0.01 m <br />) +
dry +  and fleshy +
spikelike +
8 cm80 mm <br />0.08 m <br /> (?) +  and 28 cm280 mm <br />0.28 m <br /> (?) +
subsessile +  and sessile +
bisexual +  and sterile +
terete +  and compressed +
1 cm10 mm <br />0.01 m <br /> (1.6 cm16 mm <br />0.016 m <br />) +
1 +  and 3 +
simple +  and compound +
2 (?) +  and 3 (?) +
not papillate +
Gramineae +
Elymus diversiglumis +
species +
membranous +
0 cm0 mm <br />0 m <br /> (0.02 cm0.2 mm <br />2.0e-4 m <br />) +
inconspicuous +
unequal +
plant +  and cespitose +
aquatic +  and terrestrial +