Elymus lanceolatus

(Scribn. & J.G. Sm.) Gould
Synonyms: Agropyron dasystachyum var. riparium Agropyron dasystachyum
Treatment appears in FNA Volume 24. Treatment on page 327.

Plants strongly rhizomatous, sometimes glaucous. Culms 22-130 cm, erect; nodes glabrous. Leaves often mostly basal, sometimes more evenly distributed; sheaths glabrous or pubescent; auricles usually present on the lower leaves, 0.5-1.5 mm; ligules 0.1-0.5 mm, erose, sometimes ciliolate; blades 1.5-6 mm wide, generally involute, abaxial surfaces usually glabrous, adaxial surfaces strigose, ribs subequal in size and spacing. Spikes 3.5-26 cm long, 0.5-1 cm wide, erect to slightly nodding, usually with 1 spikelet per node, sometimes with 2 at a few nodes; internodes 3.5-15 mm long, 0.1-0.8 mm wide, glabrous or hairy. Spikelets 8-31 mm, 1.5-3 times longer than the internodes, appressed, with 3-11 florets; rachillas glabrous or hairy, hairs to 1 mm; disarticulation above the glumes, beneath each floret. Glumes subequal, 5-14 mm long, 1/2 - 3/4 the length of the adjacent lemmas, 0.7-1.3 mm wide, lanceolate, glabrous or hairy, smooth or scabrous, 3-5-veined, flat or weakly, often asymmetrically keeled, keels straight, margins narrow, tapering from the base or from beyond midlength, apices acute to acuminate, sometimes mucronate or shortly awned; lemmas 7-12 mm, glabrous or hairy, hairs all alike, sometimes scabrous, acute to awn-tipped, awns to 2 mm, straight; paleas about equal to the lemmas, keels straight below the apices, smooth or scabrous proximally, sometimes hairy, scabrous distally, intercostal region glabrous or with hairs, apices 0.2-0.3 mm wide; anthers (2.5) 3-6 mm. 2n = 28.

Distribution

Oreg., Nev., Colo., N.Mex., Alaska, Ill., Mont., Wyo., Mich., Wis., Idaho, Alta., B.C., Man., Ont., Sask., Yukon, Wash., Ariz., Calif., N.Dak., Nebr., S.Dak.

Discussion

Elymus lanceolatus grows in sand and clay soils and dry to mesic habitats. It is found primarily in the western half of the Flora region, between the coastal mountains and 95° W longitude, with the exception of E. lanceolatus subsp. psammophilus, which extends around the Great Lakes. Three subspecies are recognized, primarily on the basis of their lemma and palea pubescence.

Elymus lanceolatus is primarily outcrossing, and hybridizes with several species of Triticeae. Elymus albicans (p. 334) is thought to be derived from hybridization with the awned phase of Pseudoroegneria spicata (p. 281). Judging from specimens of controlled hybrids, hybridization with E. trachycaulus (p. 321) and unawned plants of P. spicata probably occur, but would be almost impossible to detect without careful observation in the field. Experimental hybrids are partially fertile, and capable of backcrossing to either parent (Dewey 1965, 1967, 1968, 1975, 1976).

Selected References

None.

Key

1 Lemmas densely hairy, hairs flexible, some 1 mm long or longer Elymus lanceolatus subsp. psammophilus
1 Lemmas glabrous or with stiff hairs shorter than 1 mm. > 2
2 Lemmas with hairs, not scabrous Elymus lanceolatus subsp. lanceolatus
2 Lemmas smooth, sometimes scabrous distally, mosdy glabrous, sometimes the lemma margins hairy proximally Elymus lanceolatus subsp. riparius

"decumbent" is not a number.

... more about "Elymus lanceolatus"
membranous +  and scarious +
with hairs +  and scabrous +
2.5 cm25 mm <br />0.025 m <br /> (?) +
0.3 cm3 mm <br />0.003 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
bidentate +  and acute +
0.2mm;0.3mm +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
Mary E. Barkworth +, Julian J.N. Campbell +  and Bjorn Salomon +
(Scribn. & J.G. Sm.) Gould +
1 (?) +  and 3 (?) +
straight +
basal +  and apical +
from the sinus +  and terminal +
not geniculate +
0 cm0 mm <br />0 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
keeled +  and rounded +
pseudopetiolate +  and branching +
intravaginal +, extravaginal +, branching +  and basal +
swelling +
not evident +
cross +, linear +  and narrowly lanceolate +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
spikelike +
0 cm0 mm <br />0 m <br /> (1 cm10 mm <br />0.01 m <br />) +
hairy +  and glabrous +
not branching +
sometimes strongly decumbent +  and prostrate +
22 cm220 mm <br />0.22 m <br /> (130 cm1,300 mm <br />1.3 m <br />) +
not woody +
Oreg. +, Nev. +, Colo. +, N.Mex. +, Alaska +, Ill. +, Mont. +, Wyo. +, Mich. +, Wis. +, Idaho +, Alta. +, B.C. +, Man. +, Ont. +, Sask. +, Yukon +, Wash. +, Ariz. +, Calif. +, N.Dak. +, Nebr. +  and S.Dak. +
liquid +, soft +  and hard +
capillary +
pistillate +  and staminate +
3 +  and 11 +
laterally compressed +  and terete +
shorter than to longer than the adjacent florets +
5mm +  and 14mm +
subtending +
hairy +  and glabrous +
1/2 +  and 3/4 +
keeled +  and lanceolate +
subequal +
0.7mm +  and 1.3mm +
0 cm0 mm <br />0 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
with hairs +  and glabrous +
solid +  and hollow +
1.5 cm15 mm <br />0.015 m <br /> (?) +
3.5mm +  and 15mm +
hairy +  and glabrous +
0.2 cm2 mm <br />0.002 m <br /> (2.6 cm26 mm <br />0.026 m <br />) +
0.01 cm0.1 mm <br />1.0e-4 m <br /> (?) +  and 0.08 cm0.8 mm <br />8.0e-4 m <br /> (?) +
hairy +, scabrous +  and smooth +
distributed +
hairy +  and glabrous +
linear-lanceolate +
0.7 cm7 mm <br />0.007 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
stiffly membranous +  and coriaceous +
erose +  and entire +
acute +, rounded +  and truncate +
0.01 cm0.1 mm <br />1.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
membranous +
inconspicuous +
lanceolate +
fleshy +  and membranous +
tapering +
concealed +  and prominent +
few +  and 2 +
2 +  and 1 +
hairy +  and glabrous +
well-developed +
slightly longer +
subequal +
dry +  and fleshy +
spikelike +
hairy +  and glabrous +
pubescent +  and glabrous +
3.5 cm35 mm <br />0.035 m <br /> (?) +  and 26 cm260 mm <br />0.26 m <br /> (?) +
subsessile +  and sessile +
1.5-3 times longer than +
bisexual +  and sterile +
terete +  and compressed +
0.8 cm8 mm <br />0.008 m <br /> (3.1 cm31 mm <br />0.031 m <br />) +
1 +  and 3 +
simple +  and compound +
2 (?) +  and 3 (?) +
not papillate +
Agropyron dasystachyum var. riparium +  and Agropyron dasystachyum +
Elymus lanceolatus +
species +
membranous +
0 cm0 mm <br />0 m <br /> (0.02 cm0.2 mm <br />2.0e-4 m <br />) +
inconspicuous +
unequal +
aquatic +  and terrestrial +