familyPoaceae
subfamilyPoaceae subfam. Pooideae
tribePoaceae tribe Triticeae
genusElymus
speciesElymus lanceolatus
subspeciesElymus lanceolatus subsp. psammophilus

Elymus lanceolatus subsp. psammophilus

(J.M. Gillett & H. Senn) Á. Löve
Mary E. Barkworth, Julian J.N. Campbell, Bjorn Salomon
Common names: Sand-dune wheatgrass
Synonyms: Agropyron dasystachyum var. psammophilum Elymus calderi Agropyron psammophilum
Treatment appears in FNA Volume 24. Treatment on page 327.
Please click on the illustration for a higher resolution version.
Illustrator: Cindy Roché, Annaliese Miller

Copyright: Utah State University

Culms 20-95 cm. Spikes 4-26 cm; internodes 3.5-13 mm, hairy at least distally. Spikelets 9-31 mm. Lemmas densely hairy, hairs flexible, usually many longer than 1 mm; paleas hairy between the keels, keels hairy proximally.

Discussion

Elymus lanceolatus subsp. psammophilus tends to grow in sandy soils. It was originally described from around the Great Lakes, but plants with similar vestiture have been found scattered throughout the western range of the species, almost always in association with sandy soils. Those from the Yukon and northern British Columbia tend to be shorter and have smaller spikelets and spikelet parts than those from Washington and Saskatchewan, but there is con¬siderable overlap in these characters. Plants from around the Great Lakes (Gillett and Senn 1960) were almost completely pollen sterile. Despite this, Gillett and Senn rejected the notion that they were hybrids.

Selected References

None.

Lower Taxa

None.

"decumbent" is not a number.

... more about "Elymus lanceolatus subsp. psammophilus"
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2n chromosome quantity
28 +
Abaxial ligule presence
absent +
Abaxial ligule quantity
common +
Abaxial surface architecture or pubescence or relief
smooth +
Abaxial surface pubescence
glabrous +
Abaxial surface pubescence or relief
scabrous +
Adaxial cell presence
absent +
Adaxial cell shape
bulliform +
Adaxial ligule architecture or pubescence or shape
ciliate +
Adaxial ligule presence
absent +
Adaxial ligule texture
membranous +  and scarious +
Adaxial surface pubescence
strigose +
Adaxial surface pubescence or relief
with hairs +  and scabrous +
Anther atypical some measurement
2.5 cm25 mm <br />0.025 m <br /> (?) +
Anther quantity
3 +
Anther some measurement
0.3 cm3 mm <br />0.003 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
Apex architecture or shape
awned +
Apex pubescence
hairy +
Apex shape
bidentate +  and acute +
Apex width
0.2mm;0.3mm +
Arm cell presence
absent +
Auricle presence
absent +
Auricle some measurement
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
Author
Mary E. Barkworth +, Julian J.N. Campbell +  and Bjorn Salomon +
Authority
(J.M. Gillett & H. Senn) Á. Löve +
Awn architecture
unbranched +
Awn arrangement
divergent +
Awn atypical quantity
1 (?) +  and 3 (?) +
Awn course
straight +
Awn orientation
arising +
Awn position
basal +  and apical +
Awn position or structure subtype
from the sinus +  and terminal +
Awn quantity
single +
Awn shape
not geniculate +
Awn some measurement
0 cm0 mm <br />0 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
Back shape
keeled +  and rounded +
Basal floret quantity
1 +  and 2 +
Base architecture
pseudopetiolate +  and branching +
Base arrangement
close +
Base fusion
free +
Base position
intravaginal +, extravaginal +, branching +  and basal +
Base size
swelling +
Base texture
hard +
Base width
thick +
Blade arrangement
parallel +
Blade arrangement or course or shape
linear +
Blade prominence
not evident +
Blade shape
cross +, linear +  and narrowly lanceolate +
Blade shape or vernation
involute +
Blade width
0.15 cm1.5 mm <br />0.0015 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
Branch shape
spikelike +
Bristle some measurement
0 cm0 mm <br />0 m <br /> (1 cm10 mm <br />0.01 m <br />) +
Callus pubescence
hairy +  and glabrous +
Common name
Sand-dune wheatgrass +
Culm architecture
not branching +
Culm architecture or arrangement or growth form
solitary +
Culm atypical some measurement
220 +
Culm duration
annual +
Culm growth form
climbing +
Culm growth form or location
floating +
Culm growth form or orientation
sometimes strongly decumbent +  and prostrate +
Culm orientation
erect +
Culm pubescence
glabrous +
Culm some measurement
20 cm200 mm <br />0.2 m <br /> (95 cm950 mm <br />0.95 m <br />) +
Culm texture
not woody +
Distal floret reproduction
sterile +
Distal floret size
reduced +
Distribution
Oreg. +, Nev. +, Colo. +, N.Mex. +, Alaska +, Ill. +, Mont. +, Wyo. +, Mich. +, Wis. +, Idaho +, Alta. +, B.C. +, Man. +, Ont. +, Sask. +, Yukon +, Wash. +, Ariz. +, Calif. +, N.Dak. +, Nebr. +  and S.Dak. +
Embryo quantity
1/3 +
Endosperm texture
liquid +, soft +  and hard +
Filament shape
capillary +
Floret architecture
pistillate +  and staminate +
Floret arrangement
adjacent +
Floret length or size
longer +
Floret quantity
3 +  and 11 +
Floret reproduction
bisexual +
Floret shape
laterally compressed +  and terete +
Glume architecture
3-5-veined +
Glume architecture or pubescence or relief
smooth +
Glume architecture or shape
awned +
Glume height or length or size
shorter than to longer than the adjacent florets +
Glume length
5mm +  and 14mm +
Glume position
subtending +
Glume presence
absent +
Glume pubescence
hairy +  and glabrous +
Glume pubescence or relief
scabrous +
Glume quantity
1/2 +  and 3/4 +
Glume shape
keeled +  and lanceolate +
Glume size
subequal +
Glume variability
equal +
Glume width
0.7mm +  and 1.3mm +
Hair architecture
awn-tipped +
Hair fragility
pliable +
Hair pubescence or relief
scabrous +
Hair quantity
many +
Hair shape
acute +
Hair some measurement
1 cm10 mm <br />0.01 m <br /> (?) +
Hair variability
alike +
Hilum arrangement or course or shape
linear +
Hilum coloration or relief
punctate +
Hilum shape
Ovate (?) +
Illustration
Present +
Illustration copyright
Utah State University +
Illustrator
Cindy Roché +  and Annaliese Miller +
Inflorescence architecture
compound +
Inflorescence position or structure subtype
terminal +
Intercostal region pubescence
with hairs +  and glabrous +
Internode architecture
solid +  and hollow +
Internode atypical some measurement
1.5 cm15 mm <br />0.015 m <br /> (?) +
Internode length
3.5mm +  and 15mm +
Internode presence
absent +
Internode pubescence
hairy +
Internode some measurement
0.35 cm3.5 mm <br />0.0035 m <br /> (1.3 cm13 mm <br />0.013 m <br />) +
Internode width
0.01 cm0.1 mm <br />1.0e-4 m <br /> (?) +  and 0.08 cm0.8 mm <br />8.0e-4 m <br /> (?) +
Keel pubescence
hairy +
Keel pubescence or relief
scabrous +
Layer presence
absent +
Leaf arrangement
distributed +
Leaf arrangement or density
concentrated +
Leaf margin arrangement
Overlapping (?) +
Leaf position
basal +
Lemma architecture
5(7)-veined +
Lemma architecture or shape
not +  and awned +
Lemma arrangement
adjacent +
Lemma height or length or size
shorter +
Lemma length or size
longer +
Lemma pubescence
hairy +
Lemma shape
linear-lanceolate +
Lemma some measurement
0.7 cm7 mm <br />0.007 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
Lemma texture
stiffly membranous +  and coriaceous +
Ligule architecture
erose +  and entire +
Ligule architecture or relief
erose +
Ligule presence
absent +
Ligule pubescence
ciliolate +
Ligule shape
acute +, rounded +  and truncate +
Ligule some measurement
0.01 cm0.1 mm <br />1.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
Ligule texture
membranous +
Lodicule architecture
veined +
Lodicule architecture or pubescence or shape
ciliate +
Lodicule architecture or shape
truncate +
Lodicule prominence
inconspicuous +
Lodicule quantity
2 +
Lodicule shape
lanceolate +
Lodicule texture
fleshy +  and membranous +
Lowest floret architecture or shape
glumelike +
Lowest floret function
functional +
Lowest floret reproduction
sterile +
Margin shape
tapering +
Margin size or width
narrow +
Midrib architecture
simple +
Node prominence
concealed +  and prominent +
Node pubescence
glabrous +
Node quantity
few +  and 2 +
Number of lower taxa
0 +
Other quantity
2 +  and 1 +
Ovary presence
absent +
Ovary pubescence
hairy +  and glabrous +
Palea development
well-developed +
Palea height or length or size
shorter +
Palea length or size
slightly longer +
Palea presence
absent +
Palea pubescence
hairy +
Palea shape
keeled +
Palea size
subequal +
Palea variability
equal +
Papilla presence
absent +
Pedicel presence
absent +
Pedicel quantity
0-4 mm +
Pericarp arrangement
separating +
Pericarp condition or texture
dry +
Pericarp texture
dry +  and fleshy +
Pericarp width
thin +
Pseudopetiole presence
absent +
Raceme shape
spikelike +
Rachilla length
prolonged +
Rachilla pubescence
hairy +  and glabrous +
Rachis prominence
evident +
Reference
None +
Sheath architecture
open +
Sheath condition
closed +
Sheath length
open +
Sheath pubescence
pubescent +  and glabrous +
Source xml
https://bibilujan@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/84153f6d59a0a91d69695978a64cee7560374f8e/V24/V24 463.xml +
Spike architecture
ebracteate +
Spike length
3.5 cm35 mm <br />0.035 m <br /> (?) +  and 26 cm260 mm <br />0.26 m <br /> (?) +
Spike some measurement
4 cm40 mm <br />0.04 m <br /> (26 cm260 mm <br />0.26 m <br />) +
Spikelet architecture
subsessile +  and sessile +
Spikelet arrangement
divergent +
Spikelet atypical quantity
5 (?) +
Spikelet fixation
attached +
Spikelet fixation or orientation
appressed +
Spikelet fusion
distinct +
Spikelet length or size
1.5-3 times longer than +
Spikelet orientation
Patent (Usually appressed) +
Spikelet quantity
1 +
Spikelet reproduction
bisexual +  and sterile +
Spikelet shape
terete +  and compressed +
Spikelet some measurement
0.9 cm9 mm <br />0.009 m <br /> (3.1 cm31 mm <br />0.031 m <br />) +
Stamen quantity
1 +  and 3 +
Starch-grain architecture
simple +  and compound +
Style-branch atypical quantity
4 +  and 1 +
Style-branch quantity
2 (?) +  and 3 (?) +
Style quantity
2 +
Subsidiary-cell architecture
parallel-sided +
Surface relief
not papillate +
Synonyms
Agropyron dasystachyum var. psammophilum +, Elymus calderi +  and Agropyron psammophilum +
Taxon family
Poaceae +
Taxon name
Elymus lanceolatus subsp. psammophilus +
Taxon parent
Elymus lanceolatus +
Taxon rank
subspecies +
Terminal floret reproduction
sexual +
Tissue coloration
hyaline +
Tissue texture
membranous +
Tooth some measurement
0 cm0 mm <br />0 m <br /> (0.02 cm0.2 mm <br />2.0e-4 m <br />) +
Vein architecture or shape
ribbed +
Vein arrangement
spaced +
Vein presence
absent +
Vein prominence
inconspicuous +
Vein quantity
1 (?) +
Vein size
unequal +
Volume
Volume 24 +
Whole organism architecture
rhizomatous +
Whole organism arrangement or pubescence
tufted +
Whole organism duration
perennial +
Whole organism growth form
plant +
Whole organism growth form or habitat
aquatic +  and terrestrial +
Whole organism pubescence
glaucous +
X chromosome quantity
7 +