Epilobium densiflorum

(Lindley) Hoch & P. H. Raven

Phytologia 73: 457. 1993.

Common names: Denseflower willowherb
Basionym: Oenothera densiflora Lindley Edwards’s Bot. Reg. 19: plate 1593. 1833
Synonyms: Boisduvalia bipartita Greene B. densiflora (Lindley) Bartling B. densiflora var. bipartita (Greene) Jepson B. densiflora var. imbricata Greene B. densiflora var. montana Jepson B. densiflora var. pallescens Suksdorf B. densiflora var. salicina (Rydberg) Munz B. imbricata (Greene) A. Heller B. salicina Rydberg B. sparsiflora A. Heller B. sparsifolia A. Nelson & P. B. Kennedy O. densiflora var. imbricata (Greene) H. Léveillé
Treatment appears in FNA Volume 10.

Herbs usually with taproot, sometimes with loose network of roots. Stems erect or ascending, terete, 4–150 cm, simple or branched with strong central axis, proximal branches ascending or suberect, villous or strigillose, often mixed glandular puberulent distally. Leaves opposite and often early-deciduous proximally, alternate and crowded distally, usually subsessile, rarely petiole 1–2 mm, blade usually narrowly lanceolate to sublinear, rarely to lanceolate, 1.4–7.5 (–9.2) × 0.5–1.4 cm, base cuneate to attenuate, margins remotely to sharply serrulate, 5–12 teeth per side, lateral-veins obscure, 2–5 per side, apex acute, surfaces densely villous and/or strigillose; bracts broader than cauline leaves, broadly lanceolate to ovate or subrotund, 0.5–2.5 × 0.3–1.8 cm, long-acuminate, sometimes folded on midrib. Inflorescences erect spikes, congested, simple, densely villous and strigillose, sometimes mixed glandular puberulent. Flowers erect, often hidden within subtending bracts, usually chasmogamous; buds sessile, narrowly elongate, 2–4 mm; floral-tube 1.3–3.8 × 1–2.2 mm, ring of hairs 0.6–2 mm distal to base inside; sepals 2–7.5 × 0.5–2.2 mm, apex acute; petals rose-purple, magenta, pink, or white, 3–9.5 (–11.5) × 1.2–5 (–6.2) mm, apical notch 0.8–3.8 mm; filaments dark-pink, those of longer stamens 1.5–4.5 mm, those of shorter ones 0.5–1.9 mm; anthers yellow, 0.5–1.2 × 0.3–0.7 mm; ovary 2–5 mm, densely villous, often mixed glandular puberulent; style white, 2.2–5.5 (–7.5) mm, glabrous, stigma subcapitate to irregularly 4-lobed, 0.3–0.8 × 0.3–1 mm, surrounded by longer anthers. Capsules cylindrical to subfusiform, 4–11 mm, beak to 0.5 mm, central column persistent, surfaces densely villous; subsessile or pedicel 1–2.5 mm. Seeds 3–8 per locule, irregularly angular-fusiform, 1.2–1.6 (–1.9) × 0.4–1 mm, without a chalazal collar, light-brown, surface irregularly reticulate with raised cells. 2n = 20.


Phenology: Flowering May–Oct.
Habitat: Vernally wet places, moist pastures, woodlands, meadows, along streams and ditches, alluvial valleys, often on low ground in volcanic or sandy soils.
Elevation: 0–2600 m.

Distribution

B.C., Ariz., Calif., Idaho, Mont., Nev., Oreg., Utah, Wash., Mexico (Baja California)

Discussion

Epilobium densiflorum is an extremely variable species that changes its aspect through the flowering season. Collections made very early in the season may include only well-spaced, narrowly lanceolate leaves, the proximal ones usually opposite, and a short, sparse, somewhat open inflorescence. A late-season collection, even from the same population, may entirely lack cauline leaves, and consist instead of bare, peeling stems topped by dense, tightly imbricate-bracted inflorescences, with each broad bract enclosing a capsule or flower.

Boisduvalia douglasii Spach is an illegitimate substitute for Oenothera densiflora Lindley and pertains here.

Selected References

None.

Lower Taxa

None.
... more about "Epilobium densiflorum"
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.12 cm1.2 mm <br />0.0012 m <br />) +
episepalous +
subequal +  and longer +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.07 cm0.7 mm <br />7.0e-4 m <br />) +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.38 cm3.8 mm <br />0.0038 m <br />) +
Peter C. Hoch +
(Lindley) Hoch & P. H. Raven +
cuneate +  and attenuate +
Oenothera densiflora +
0 cm0 mm <br />0 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
indehiscent +  and loculicidal +
7.5 cm75 mm <br />0.075 m <br /> (9.2 cm92 mm <br />0.092 m <br />) +
1.4 cm14 mm <br />0.014 m <br /> (7.5 cm75 mm <br />0.075 m <br />) +
toothed +  and entire +
lanceolate +, usually narrowly lanceolate +  and sublinear +
0.5 cm5 mm <br />0.005 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
long-acuminate +, broadly lanceolate +  and ovate or subrotund +
reduced +
0.3 cm3 mm <br />0.003 m <br /> (?) +  and 1.8 cm18 mm <br />0.018 m <br /> (?) +
recurved +  and erect +
elongate +
0.2 cm2 mm <br />0.002 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
sessile +  and pedicellate +
curved +  and straight +
cylindrical +  and subfusiform +
0.4 cm4 mm <br />0.004 m <br /> (1.1 cm11 mm <br />0.011 m <br />) +
persistent +
Denseflower willowherb +
B.C. +, Ariz. +, Calif. +, Idaho +, Mont. +, Nev. +, Oreg. +, Utah +, Wash. +  and Mexico (Baja California) +
0–2600 m. +
basifixed +  and versatile +
0.13 cm1.3 mm <br />0.0013 m <br /> (0.38 cm3.8 mm <br />0.0038 m <br />) +
1mm +  and 2.2mm +
actinomorphic +
sessile +  and pedicellate +
curved +  and straight +
4-angled +, terete +, narrowly cylindrical +  and fusiform or rarely narrowly ellipsoidal +
slender +
Vernally wet places, moist pastures, woodlands, meadows, along streams and ditches, alluvial valleys, often on low ground in volcanic or sandy soils. +
perennial +  and annual +
suffrutescent +
puberulent +, strigillose +  and villous +
crowded +, alternate +  and opposite +
early-deciduous +
proximal +, basal +  and cauline +
decurrent +
puberulent +  and villous +
0.2 cm2 mm <br />0.002 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
numerous +  and 1 +
multicellular +
0.1 cm1 mm <br />0.001 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
0.95 cm9.5 mm <br />0.0095 m <br /> (1.15 cm11.5 mm <br />0.0115 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (0.62 cm6.2 mm <br />0.0062 m <br />) +
white +, pink +, magenta +  and rose-purple +
3mm +  and 9.5mm +
obovate +  and obcordate +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
Flowering May–Oct. +
suberect +  and ascending +
puberulent +, strigillose +  and villous +
inflated +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
basal +  and cauline +
single +, 8 +  and 12 +
clavate +  and globose +
0.16 cm1.6 mm <br />0.0016 m <br /> (0.19 cm1.9 mm <br />0.0019 m <br />) +
light-brown +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.16 cm1.6 mm <br />0.0016 m <br />) +
3 +  and 8 +
sculptured +  and smooth +
angular-fusiform +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
lanceolate +, , +, flushed with red or flushed with cream +  and green +
deciduous +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.22 cm2.2 mm <br />0.0022 m <br />) +
parenchymatous +
parietal +, axile +  and placentation +
unequal +
aquatic +, amphibious +  and terrestrial +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.19 cm1.9 mm <br />0.0019 m <br />) +
2 times as many as sepals +
branched +  and simple +
prostrate +  and decumbent +
ascending +  and erect +
glabrous +, villous +, glandular-puberulent +  and strigillose +
exfoliating +
4 cm40 mm <br />0.04 m <br /> (150 cm1,500 mm <br />1.5 m <br />) +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
subcapitate +  and irregularly 4-lobed +
0.03 cm0.3 mm <br />3.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
subsessile +  and petiolate +
intrapetiolar +
0.55 cm5.5 mm <br />0.0055 m <br /> (0.75 cm7.5 mm <br />0.0075 m <br />) +
0.22 cm2.2 mm <br />0.0022 m <br /> (0.55 cm5.5 mm <br />0.0055 m <br />) +
aquatic +, amphibious +  and terrestrial +
villous +  and strigillose +
papillose +  and finely reticulate +
constricted +
Boisduvalia bipartita +, B. densiflora +, B. densiflora var. bipartita +, B. densiflora var. imbricata +, B. densiflora var. montana +, B. densiflora var. pallescens +, B. densiflora var. salicina +, B. imbricata +, B. salicina +, B. sparsiflora +, B. sparsifolia +  and O. densiflora var. imbricata +
Epilobium densiflorum +
Epilobium sect. Pachydium +
species +
3(-5)-aperturate +