Epilobium pallidum

(Eastwood) Hoch & P. H. Raven

Phytologia 73: 458. 1993.

Common names: Largeflower spike-primrose
Basionym: Boisduvalia pallida Eastwood Leafl. W. Bot. 2: 54. 1937
Synonyms: B. macrantha A. Heller 1905
Treatment appears in FNA Volume 10.
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Herbs with taproot. Stems terete, 4.5–55 cm, simple or with proximal ascending branches and/or distal branches, ± densely villous and/or strigillose, mixed glandular puberulent distally or glabrescent. Leaves opposite only in proximal pairs, alternate and crowded distally, subsessile, blade narrowly elliptic or lanceolate, 1.2–5 × 0.2–1.2 cm, usually longer than internodes, base cuneate to attenuate, margins subentire or sparsely serrulate, 3–5 teeth per side, veins inconspicuous, 2–7 per side, apex acute, surfaces villous; bracts same size and shape as leaves. Inflorescences erect spikes, often congested proximally, sometimes nodding in bud, simple, or sometimes sparsely branched, ± densely mixed villous, strigillose, and glandular puberulent. Flowers erect, mostly chasmogamous, sometimes initiating at second or third proximal node; buds 3–5 × 2.5–3.5 mm, apex blunt; floral-tube 1.5–3 × 1.1–2.8 mm, ring of lax hairs near base inside; sepals erect, (1.5–) 2–6.2 × (0.5–) 1–2.2 mm, densely villous and glandular-pubescent abaxially; petals rose-purple, (2.8–) 4.2–9.8 × (2.2–) 3.5–5.3 mm, apical notch 1.2–3.5 mm; filaments rose-purple, those of longer stamens 2.4–6.5 mm, those of shorter ones 0.9–4.3 mm; anthers 0.6–1.6 × 0.2–0.8 mm; ovary 3.5–9 mm, densely pubescent; style pink, 3.3–9.2 mm, stigma 4-lobed, ± subentire in smaller flowers, 0.7–1.4 × 0.5–1.3 mm, usually surrounded by, rarely exserted beyond, longer anthers. Capsules narrowly fusiform, (10–) 14–21 × 1–2.1 mm, beak 2–5 mm, ribs usually broad and prominent, septa and central column disintegrating with age, surfaces ± densely villous; subsessile. Seeds 8–12 per capsule, initially in 4 locules, at maturity pushed into 2 rows proximally, 1 overlap­ping row distally, irregularly angular-fusiform, 1.8–2.3× 0.7–0.9 mm, chalazal collar absent, surface irreg­ularly reticulate. 2n = 20.


Phenology: Flowering May–Aug(–Sep).
Habitat: Stream banks and washes in vernally moist areas in shrubland or lower forested regions.
Elevation: 60–2100 m.

Discussion

P. H. Raven and D. M. Moore (1965) observed that smaller-flowered individuals of this species were sometimes difficult to separate from Epilobium torreyi (reported as Boisduvalia stricta), especially in the absence of mature fruits. The capsules of E. pallidum are usually longer and thinner than those of related species and the internal structure is highly distinctive. The stigmas of larger-flowered plants often are exserted beyond the anthers, increasing the chances for outcrossing.

Selected References

None.

Lower Taxa

None.
... more about "Epilobium pallidum"
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.16 cm1.6 mm <br />0.0016 m <br />) +
episepalous +
subequal +  and longer +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
blunt +  and acute +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
Peter C. Hoch +
(Eastwood) Hoch & P. H. Raven +
cuneate +  and attenuate +
Boisduvalia pallida +
0.2 cm2 mm <br />0.002 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
indehiscent +  and loculicidal +
1.2 cm12 mm <br />0.012 m <br /> (5 cm50 mm <br />0.05 m <br />) +
usually longer +
toothed +  and entire +
lanceolate +  and elliptic +
0.2 cm2 mm <br />0.002 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
reduced +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
recurved +  and erect +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
sessile +  and pedicellate +
1 cm10 mm <br />0.01 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
curved +  and straight +
1.4 cm14 mm <br />0.014 m <br /> (2.1 cm21 mm <br />0.021 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.21 cm2.1 mm <br />0.0021 m <br />) +
not +  and persistent +
persistent +
Largeflower spike-primrose +
Calif. +, Idaho +  and Oreg. +
60–2100 m. +
basifixed +  and versatile +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
1.1mm +  and 2.8mm +
actinomorphic +
sessile +  and pedicellate +
curved +  and straight +
4-angled +, terete +, narrowly cylindrical +  and fusiform or rarely narrowly ellipsoidal +
slender +
Stream banks and washes in vernally moist areas in shrubland or lower forested regions. +
perennial +  and annual +
suffrutescent +
branched +  and simple +
puberulent +, strigillose +  and villous +
crowded +, alternate +  and opposite +
deciduous +
proximal +, basal +  and cauline +
decurrent +
serrulate +  and subentire +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
numerous +  and 1 +
multicellular +
0.28 cm2.8 mm <br />0.0028 m <br /> (0.42 cm4.2 mm <br />0.0042 m <br />) +
0.22 cm2.2 mm <br />0.0022 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
rose-purple +
4.2mm +  and 9.8mm +
obovate +  and obcordate +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.53 cm5.3 mm <br />0.0053 m <br />) +
Flowering May–Aug(–Sep). +
prominent +
inflated +
basal +  and cauline +
clavate +  and globose +
0.18 cm1.8 mm <br />0.0018 m <br /> (0.23 cm2.3 mm <br />0.0023 m <br />) +
1 +, 8 +  and 12 +
sculptured +  and smooth +
angular-fusiform +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.09 cm0.9 mm <br />9.0e-4 m <br />) +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
lanceolate +, , +, flushed with red or flushed with cream +  and green +
deciduous +
glandular-pubescent +  and villous +
0.1 cm1 mm <br />0.001 m <br /> (0.22 cm2.2 mm <br />0.0022 m <br />) +
parenchymatous +
parietal +, axile +  and placentation +
disintegrating +
unequal +
aquatic +, amphibious +  and terrestrial +
0.09 cm0.9 mm <br />9.0e-4 m <br /> (0.43 cm4.3 mm <br />0.0043 m <br />) +
2 times as many as sepals +
with proximal ascending branches +  and simple +
prostrate +  and decumbent +
erect +  and ascending or decumbent +
glabrescent +, puberulent +, strigillose +  and villous +
exfoliating +
4.5 cm45 mm <br />0.045 m <br /> (55 cm550 mm <br />0.55 m <br />) +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.14 cm1.4 mm <br />0.0014 m <br />) +
subentire +  and 4-lobed +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.13 cm1.3 mm <br />0.0013 m <br />) +
subsessile +  and petiolate +
intrapetiolar +
0.33 cm3.3 mm <br />0.0033 m <br /> (0.92 cm9.2 mm <br />0.0092 m <br />) +
aquatic +, amphibious +  and terrestrial +
papillose +  and finely reticulate +
constricted +
B. macrantha +
Epilobium pallidum +
Epilobium sect. Pachydium +
species +
3(-5)-aperturate +
inconspicuous +