Eremothera boothii

(Douglas) W. L. Wagner & Hoch

Syst. Bot. Monogr. 83: 209. 2007.

Basionym: Oenothera boothii Douglas in W. J. Hooker Fl. Bor.-Amer. 1: 213. 1832
Synonyms: Camissonia boothii (Douglas) P. H. Raven Sphaerostigma boothii (Douglas) Walpers
Treatment appears in FNA Volume 10.

Herbs slender or stout, glabrate to strigillose or villous, and/or glandular puberulent, especially in inflorescence. Stems usually well branched at base and distally, 3–65 cm, usually flowering only distally. Leaves cauline, sometimes with lower ones clustered near base, these often withered by flowering, 1–11 (–13) × 0.2–2.2 (–3) cm; petiole 0–6 cm; blade very narrowly elliptic to narrowly ovate, lanceolate, or oblanceolate, margins denticulate, serrulate, sinuate-toothed, or subentire. Inflorescences nodding. Flowers opening at sunset; floral-tube (2–) 3–8 mm, villous in proximal 1/2 inside; sepals (2.7–) 4–8 mm; petals usually white fading pink, rarely red and fading red, (3–) 3.5–9 mm; episepalous filaments (1.5–) 2–5.8 mm, epipetalous filaments slightly shorter, anthers (1–) 1.8–2.3 mm; style (6.5–) 8.2–13.5 (–15) mm, villous near base, stigma 1.2–2 mm diam., exserted beyond anthers at anthesis. Capsules cylindrical and thickened proximally, spreading to curved downward, contorted to straight, terete or 4-angled, 8–35 × 0.9–3.8 mm, held on dried plants and regularly but tardily dehiscent. Seeds usually dimorphic, rarely monomorphic, those with relatively smooth surfaces light-brown, 1.4–2.1 × 0.5–0.7 mm, those of lower portion dark-brown, 1.4–2.1 × 0.6–0.9 mm, coarsely papillose. 2n = 14.

Distribution

w United States, nw Mexico

Discussion

Subspecies 6 (6 in the flora).

P. H. Raven (1969) included within Eremothera boothii a complex of intergrading entities that have been variously treated, ranging from three species with infraspecific taxa to one species with infraspecific taxa. Raven determined E. boothii to be self-incompatible.

Selected References

None.

Key

1 Cluster of basal leaves prominent at time of flowering; plants blooming February through June (or August), strigillose and/or glandular puberulent. > 2
2 Capsules 2–3.8 mm diam. near base, thickened and indurate along angles, curved outward but not downward; inflorescences very dense. Eremothera boothii subsp. condensata
2 Capsules 1–2.3 mm diam. near base, not thickened and indurate, curved outward or downward; inflorescences ± open. > 3
3 Capsules 1.7–2.3 mm diam. near base, curved outward. Eremothera boothii subsp. decorticans
3 Capsules 1–1.6 mm diam. near base, apex often curved downward. Eremothera boothii subsp. desertorum
1 Cluster of basal leaves rarely prominent at time of flowering, leaves evenly distributed; plants blooming May through August (or September), villous and/or glandular puberulent, rarely (subsp. alyssoides) densely strigillose. > 4
4 Herbs usually strigillose, often densely so, rarely villous and/or glandular puberulent; plants flowering May through June (or August). Eremothera boothii subsp. alyssoides
4 Herbs villous and glandular puberulent; plants flowering (May to) June through August (or September). > 5
5 Herbs villous, also glandular puberulent, especially in inflorescences; stems usually 15–60 cm; leaf blades narrowly lanceolate, narrowly ovate, or ovate to elliptic, 0.6–2.2 cm wide, margins coarsely serrulate to sinuate-toothed. Eremothera boothii subsp. boothii
5 Herbs densely villous, also densely glandular puberulent, especially in inflorescences; stems 5–20 cm; leaf blades narrowly elliptic to narrowly lanceolate or lanceolate, sometimes oblanceolate proximally, 0.3–1.5 cm wide, margins sparsely serrulate to sometimes sinuate-toothed. Eremothera boothii subsp. intermedia
... more about "Eremothera boothii"
0.1 cm1 mm <br />0.001 m <br /> (0.18 cm1.8 mm <br />0.0018 m <br />) +
0.18 cm1.8 mm <br />0.0018 m <br /> (0.23 cm2.3 mm <br />0.0023 m <br />) +
Warren L. Wagner +
(Douglas) W. L. Wagner & Hoch +
papillose +
Oenothera boothii +
indehiscent +  and loculicidal +
entire +, sinuate-toothed +, serrulate +  and denticulate +
very narrowly elliptic +  and narrowly ovate lanceolate or oblanceolate +
0.8 cm8 mm <br />0.008 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
spreading +  and curved downward +
4-angled +, terete +  and cylindrical +
0.09 cm0.9 mm <br />9.0e-4 m <br /> (0.38 cm3.8 mm <br />0.0038 m <br />) +
w United States +  and nw Mexico +
reddish green +  and white +
exfoliating +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.58 cm5.8 mm <br />0.0058 m <br />) +
basifixed +  and versatile +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
actinomorphic +
straight +
loculicidal +
4-angled +, terete +  and cylindrical +
puberulent +, glabrate +  and strigillose or villous +
stout +  and slender +
11 cm110 mm <br />0.11 m <br /> (13 cm130 mm <br />0.13 m <br />) +
2.2 cm22 mm <br />0.022 m <br /> (3 cm30 mm <br />0.03 m <br />) +
deciduous +
1 cm10 mm <br />0.01 m <br /> (11 cm110 mm <br />0.11 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (2.2 cm22 mm <br />0.022 m <br />) +
subentire +, sinuate-toothed +, serrulate +  and denticulate +
numerous +  and 1 +
0.3 cm3 mm <br />0.003 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
red and fading red +, pink +  and white fading +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
0 cm0 mm <br />0 m <br /> (6 cm60 mm <br />0.06 m <br />) +
0.14 cm1.4 mm <br />0.0014 m <br /> (0.21 cm2.1 mm <br />0.0021 m <br />) +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.09 cm0.9 mm <br />9.0e-4 m <br />) +
Syst. Bot. Monogr. +
basal +  and cauline +
clavate +  and globose +
monomorphic +
7 +  and numerous +
sculptured +  and smooth +
truncate-ellipsoid +, narrowly obovoid +  and oblanceoloid +
in pairs +  and singly +
0.27 cm2.7 mm <br />0.0027 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
purple +, pink +, red +  and green +
deciduous +
0.4 cm4 mm <br />0.004 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
parenchymatous +
parietal +, axile +  and placentation +
unequal +
aquatic +, amphibious +  and terrestrial +
episepalous +
2 times as many or as many as sepals +
prostrate +  and decumbent +
ascending +  and erect +
3 cm30 mm <br />0.03 m <br /> (65 cm650 mm <br />0.65 m <br />) +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
subglobose +
subsessile +  and petiolate +
intrapetiolar +
1.35 cm13.5 mm <br />0.0135 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
0.82 cm8.2 mm <br />0.0082 m <br /> (1.35 cm13.5 mm <br />0.0135 m <br />) +
aquatic +, amphibious +  and terrestrial +
0.14 cm1.4 mm <br />0.0014 m <br /> (?) +  and 0.21 cm2.1 mm <br />0.0021 m <br /> (?) +
non-papillate +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (?) +  and 0.07 cm0.7 mm <br />7.0e-4 m <br /> (?) +
Camissonia boothii +  and Sphaerostigma boothii +
Eremothera boothii +
Eremothera +
species +
3(-5)-aperturate +
smooth +  and erose +
papillate +