Frangula purshiana subsp. ultramafica

J. O. Sawyer & S. W. Edwards

Madroño 54: 172, fig. 1. 2007.

Common names: Caribou coffeeberry
Endemic
Treatment appears in FNA Volume 12. Treatment on page 58. Mentioned on page 57.

Plants 1–2 m, shrublike. Twigs green to gray or dull brown, densely hairy. Leaves deciduous or semideciduous; blade broadly oblong or broadly elliptic to ovate or obovate, (3.5–) 5–10 cm, distinctly coriaceous, base mostly obtuse or tapered, margins serrulate or entire, often wavy, apex obtuse, often notched, surfaces papillate, sparsely to densely hairy or adaxial velvety, glaucous adaxially when fresh.


Phenology: Flowering Apr–Jun.
Habitat: Seeps, montane chaparral, open forests over mafic and ultramafic substrates.
Elevation: 800–2000 m.

Discussion

Of the three subspecies of Frangula purshiana, subsp. ultramafica is the most distinctive and might warrant species status. It appears to be restricted to the Feather River complex of serpentinized peridotite and associated mafic and ultramafic substrates near Bucks Lake in Plumas County. Its firm, bluish or greenish gray leaves are suggestive of evergreen F. californica subsp. tomentella, but they are broader and larger and bear only simple, erect hairs. The leaves are deciduous as in F. rubra, but the large, broad leaves and fruits with three stones are like those of F. purshiana.

Selected References

None.

Lower Taxa

None.
glaucous +, velvety +  and hairy +
perigynous +  and epigynous +
notched +  and obtuse +
John O. Sawyer† +  and Guy L. Nesom +
J. O. Sawyer & S. W. Edwards +
tapered +  and obtuse +
Rhamnus purshiana +
protruding +
3.5 cm35 mm <br />0.035 m <br /> (5 cm50 mm <br />0.05 m <br />) +
green +  and bluish or greenish gray +
3[-5]-veined +  and pinnate +
spinulose +, spinose +, serrate +  and entire +
denticulate +, dentate +, crenulate +, crenate +  and serrulate +
broadly elliptic;ovate or obovate +
5 cm50 mm <br />0.05 m <br /> (10 cm100 mm <br />0.1 m <br />) +
coriaceous +
Caribou coffeeberry +
globose +  and depressed-globose +
0.5 cm5 mm <br />0.005 m <br /> (1 cm10 mm <br />0.01 m <br />) +
800–2000 m. +
Seeps, montane chaparral, open forests over mafic and ultramafic substrates. +
circumscissile +
free +  and adnate +
unisexual +  and bisexual +
10-25-flowered +  and pedunculate +
semideciduous +  and deciduous +
entire +  and serrulate +
intrastaminal +
0.5 cm5 mm <br />0.005 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
not fleshy +
perigynous +  and epigynous +
adnate +  and distinct +
clawed +, broadly obovate +  and obcordate +
0.6 cm6 mm <br />0.006 m <br /> (2.3 cm23 mm <br />0.023 m <br />) +
Flowering Apr–Jun. +
2-4-carpellate +
not furrowed +
obovoid +  and lenticular +
yellowish +  and green or white +
distinct +
spreading +  and erect +
keeled;ovate-triangular +
red +  and brown gray or green +
hairy +  and glabrous +
2-3-lobed +
indehiscent +
glabrous or +  and sparsely densely hairy +
papillate +
Rhamnus sect. Frangula +  and Rhamnus subg. Frangula +
Frangula purshiana subsp. ultramafica +
Frangula purshiana +
subspecies +
green;gray or dull brown +
deciduous +  and evergreen +
plant +  and shrublike +
polygamous +, dioecious +  and synoecious +