Geum laciniatum

Murray

Novi Comment. Soc. Regiae Sci. Gott. 5: 30, plate 2. 1775.

Common names: Rough avens benoîte lacinicée
Endemic
Synonyms: Geum laciniatum var. trichocarpum Fernald
Treatment appears in FNA Volume 9. Treatment on page 68. Mentioned on page 60, 61, 69, 70.

Plants leafy-stemmed. Stems 30–100 cm, hirsute, some hairs 2–2.5 mm. Leaves: basal 15–32 cm, blade simple, lyrate-pinnate, or pinnate, major leaflets 1–7, alternating with 0–10 minor leaflets, terminal leaflet slightly to much larger than major laterals; cauline 3.5–18 cm, stipules ± free, 4–14 × 7–17 mm, blade pinnate, 3-foliolate, or simple and unlobed. Inflorescences 2–9-flowered. Pedicels densely puberulent, hirsute, eglandular. Flowers erect; epicalyx bractlets 1–2 mm; hypanthium green; sepals spreading but soon reflexed, 3–10 mm; petals spreading, white, oblong to elliptic, 2–5 mm, shorter than sepals, apex obtuse to rounded. Fruiting tori sessile, glabrous except for ring of bristles at base and tuft at apex. Fruiting styles geniculate-jointed, proximal segment persistent, 2.5–5 mm, apex hooked, glabrous, sometimes 1–2 eglandular bristles at base, distal segment deciduous, 1–2 mm, short hairs on basal 1/2.2n = 42.


Phenology: Flowering early summer.
Habitat: Wet woods and thickets, flood plains, wet woods around lakes, stream banks, boggy meadows
Elevation: 0–1000 m

Distribution

V9 99-distribution-map.jpg

N.B., N.S., Ont., P.E.I., Que., Conn., Del., D.C., Ill., Ind., Iowa, Kans., Ky., Maine, Md., Mass., Mich., Minn., Mo., Nebr., N.H., N.J., N.Y., N.C., Ohio, Pa., R.I., S.Dak., Tenn., Vt., Va., W.Va., Wis.

Discussion

B. L. Robinson and M. L. Fernald (1908) and J. K. Small (1933) misapplied the name Geum virginianum to this species. As a consequence, older specimens of G. laciniatum are often labeled (and filed in herbaria) as G. virginianum.

Geum laciniatum is unique among members of the genus in having some of the heads of achenes pop off the stem and disperse as a unit. The heads disarticulate where the torus joins the hypanthium, leaving the hypanthium inverted at the tip of the stem.

Fernald described var. trichocarpum based on the presence of bristles on the summits of the achenes. Although this morphology occurs to the near exclusion of the glabrous one in the western part of the species range, it also occurs as far east as the Carolinas, Maine, New Jersey, and Virginia. The glabrous condition has a smaller range, and it is the prominent one in Nova Scotia, Kentucky, Maine, and Ohio. The two are well mixed in Quebec, New York, and Pennsylvania. Although they are easily distinguished (hairs absent versus present), they seem to have little phytogeographic significance, and presence of achene hairs is not correlated with any other characteristics. It appears to be a trivial variation.

Geum laciniatum hybridizes with G. urbanum (= G. ×macneillii J.-P. Bernard & R. Gauthier); see discussion under 15. G. urbanum.

Selected References

None.

Lower Taxa

None.

"thin" is not a number."elongating" is not a number. "elongating" is not a number.

... more about "Geum laciniatum"
hooked;obtuse;rounded +
Joseph R. Rohrer +
Murray +
compound +  and simple +
opposite +  and alternate +
15 cm150 mm <br />0.15 m <br /> (32 cm320 mm <br />0.32 m <br />) +
simple +, 3-foliolate +  and pinnate +
pinnate;lyrate-pinnate;pinnate;lyrate-pinnate +
unlobed +
coriaceous +
free +  and distinct +
3.5 cm35 mm <br />0.035 m <br /> (18 cm180 mm <br />0.18 m <br />) +
Rough avens +  and benoîte lacinicée +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
N.B. +, N.S. +, Ont. +, P.E.I. +, Que. +, Conn. +, Del. +, D.C. +, Ill. +, Ind. +, Iowa +, Kans. +, Ky. +, Maine +, Md. +, Mass. +, Mich. +, Minn. +, Mo. +, Nebr. +, N.H. +, N.J. +, N.Y. +, N.C. +, Ohio +, Pa. +, R.I. +, S.Dak. +, Tenn. +, Vt. +, Va. +, W.Va. +  and Wis. +
not +  and aggregated +
0–1000 m +
0.1 cm1 mm <br />0.001 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (4.6 cm46 mm <br />0.046 m <br />) +
unisexual +  and bisexual +
not +  and aggregated +
tapered +, ovoid +  and fusiform +
0.2 cm2 mm <br />0.002 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
Wet woods and thickets, flood plains, wet woods around lakes, stream banks, boggy meadows +
0.2 cm2 mm <br />0.002 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
saucer-shaped +  and cupshaped +
0.2 cm2 mm <br />0.002 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
crassinucellate +
odd-pinnate +, lyrate-pinnate +, 3-foliolate +  and simple +
opposite +  and alternate +
winter-persistent +
cauline +  and basal +
cordate +, reniform +, orbiculate +, oblanceolate +, obovate +, oblong +, elliptic +  and rhombic +
palmate +  and pinnate +
crenate +, dentate +, laciniate +, 2-7-lobed +, flat +, lobed +, toothed +  and entire +
inferior +  and superior +
clustered +, biseriate +  and superposed +
hirsute +  and puberulent +
purple-veined +
0 (?) +  and 4 (?) +
free +  and distinct +
oblong;elliptic +
0.2 cm2 mm <br />0.002 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
Flowering early summer. +
adnate +, free +, connate +  and distinct +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
Novi Comment. Soc. Regiae Sci. Gott. +
not arillate +
persistent +
free +  and distinct +
reflexed +  and spreading +
deltate-lanceolate +, deltate-ovate +  and deltate +
0.3 cm3 mm <br />0.003 m <br /> (1 cm10 mm <br />0.01 m <br />) +
free +  and distinct +
1 +  and 5 +
30 cm300 mm <br />0.3 m <br /> (100 cm1,000 mm <br />1 m <br />) +
palmate +  and pinnate +
0.4 cm4 mm <br />0.004 m <br /> (1.4 cm14 mm <br />0.014 m <br />) +
ovate;lanceolate;ovate;lanceolate;linear +
0.7 cm7 mm <br />0.007 m <br /> (1.7 cm17 mm <br />0.017 m <br />) +
geniculate-jointed +  and entire +
persistent +
distinct +
basal +, lateral +, subterminal +  and terminal +
hairy +  and glabrous +
not hooked +  and hooked +
accrescent +
10 cm100 mm <br />0.1 m <br /> (?) +
Geum laciniatum var. trichocarpum +
Geum laciniatum +
species +
slightly to much larger +
inconspicuous +
hemispheric +  and cylindric +
enlarged +
geniculate-jointed +
hairy +  and glabrous +