Grimmia subg. Guembelia

(Hampe) Schimper

Coroll. Bryol. Eur., 48. 1856,.

Basionym: Undefined subg. Guembelia Hampe Bot. Zeitung (Berlin) 4: 124. 1846
Treatment appears in FNA Volume 27. Treatment on page 233. Mentioned on page 226, 229, 241, 248, 258.

Plants 10–20 mm (–70 mm for G. atrata). Stem central strand present or absent. Leaves narrowly ovatelanceolate to oblong-lanceolate, keeled, margins plane, recurved or incurved, costa prominent; basal marginal cells hyaline or not; distal lamina 2-stratose or with at least several marginal rows 2-stratose. Gemmae absent. Sexual condition autoicous or dioicous. Seta straight, centrally attached to capsule. Capsule immersed to exserted, smooth (except G. mariniana); stomata present or absent, small, at neck to part-way up capsule. Calyptra mitrate to cucullate, covering operculum.

Distribution

North America, Mexico, Central America, South America, Eurasia, Africa, Pacific Islands, Australia, Antarctic

Discussion

Species 41 (13 in the flora).

Members of subg. Guembelia are recognized by their thick, keeled leaves and usually long, straight setae with smooth capsules. Most occur on dry acidic rock, with three species growing on damp to wet acidic rock and one on dry calcareous rock.

Selected References

None.

... more about "Grimmia subg. Guembelia"
deciduous +  and persistent +
well-defined +
differentiated +
Roxanne I. Hastings +  and Henk C. Greven +
(Hampe) Schimper +
only rarely longer +
not papillose +
toothed +
sinuous +  and straight to sometimes +
sinuose-nodulose;quadrate;elongate +
oblate;elongate +
divided +
Undefined subg. Guembelia +
undifferentiated +
plicate +  and smooth +
less +  and 1/2 +
not papillose +
mitrate +  and cucullate +
small;medium +
sulcate +  and smooth +
pendent +  and erect +
striate +  and smooth +
rarely ventricose ovoid;obloid rarely globose or cylindric +
thin +  and thick +
thick-walled +  and thin-walled +
quadrate +  and rectangular +
excurrent;percurrent;excurrent +
subpercurrent +
reniform +, elliptical +  and terete +
usually not markedly larger +
with at-least several marginal rows +  and 2-stratose +
North America +, Mexico +, Central America +, South America +, Eurasia +, Africa +, Pacific Islands +, Australia +  and Antarctic +
reproduction +  and asexual +
dioicous +, autoicous +  and sexual +
papillose +, mammillose +  and smooth +
specialized +
dioicous +, autoicous +  and sexual +
keeled +, narrowly ovatelanceolate +  and oblong-lanceolate +
1;multistratose +
incurved +  and recurved +
thick-walled +  and sinuose +
thick-walled +  and sinuose +
rostrate;conic;rostrate;conic;mammillate +
not +  and enlarged +
not +  and differentiated +
lanceolate;linear +
Coroll. Bryol. Eur., +
straight +
sigmoid +  and arcuate +
papillose +  and smooth +
short to long +
ornamented +  and smooth +
globose +
muticous;long-awned +
Dryptodon +  and Hydrogrimmia +
Grimmia subg. Guembelia +
subgenus +
dark black-green;rusty-redbrown +