Hedwigia detonsa

(M. Howe) W. R. Buck & D. H. Norris

Nova Hedwigia 62: 368. 1996.

EndemicIllustrated
Basionym: Hedwigia albicans var. detonsa M. Howe Erythea 5: 91. 1897
Treatment appears in FNA Volume 28. Treatment on page 87. Mentioned on page 84, 85, 86.

Leaves 2–2.5 mm; margins broadly or narrowly recurved in basal 1/4 or plane on one or both sides, erect to incurved in acumen, entire in apex; apex erect when dry, broad, hyaline, acumen broad-based, 1/5–2/5 (–1/2) leaf length, channeled to tubular, extreme apex flat or sometimes twisted; medial and distal laminal cells with papillae 1 (or 2), large, stalked, strongly, regularly branched, branches usually of equal lengths, especially on abaxial surface; laminal cells in hyaline area with papillae low, simple at base, in longitudinal lines on cell lumina; apical cell long-linear, sharply pointed, (80–) 110–125 (–150) µm, multipapillose with papillae minute, simple, scattered on lumina and margins, or sometimes smooth. Perichaetia with leaves broadly channeled to involute, margins entire. Vaginula sparsely pilose. Calyptra naked to sparsely pilose.


Phenology: Capsules mature spring.
Habitat: River canyons, open forested lands, dry or moist acidic rock, sandstone, boulders and cliff faces near creeks, open, diffusely lighted mixed forests of Aesculus californica, Calocedrus, Pinus coulteri, Quercus chrysolepis, Torreya, and Umbellularia
Elevation: low to moderate elevations (200-1500 m)

Discussion

Unlike the irregular papillae of Hedwigia ciliata and H. stellata, those of H. detonsa as seen in cross section are nearly identical in regularity of form, moderately stalked and evenly branched. Current studies suggest the species is endemic to California, but it may also be expected in Baja California (W. R. Buck and D. H. Norris 1996). These authors find H. detonsa characteristic of river canyons and open, forested lands, more inland than H. stellata, and avoiding grasslands and savannas.

Hedwigia nivalis (Müller Hal.) Mitten

As noted above, the occurrence of Hedwigia nivalis in the flora area was discussed by W. R. Buck and D. H. Norris (1996), but without definitive conclusion.

Hedwigia ciliata var. leucophaea Bruch & Schimper

Only the typical variety of Hedwigia ciliata was accepted for the flora of North America by L. E. Anderson et al. (1990). Variety leucophaea is an established European taxon accepted as a northern variant of H. ciliata by L. Hedenäs (1994), but it was not reported or recognized in several northern or circumboreal floras, such as that of Canada (R. R. Ireland et al. 1987).

Selected References

None.

Lower Taxa

None.

"narrow" is not a number. "wide" is not a number."channeled" is not a number. "tubular" is not a number."wide" is not a number.

... more about "Hedwigia detonsa"
broad-based +
canaliculate-furrowed +
papillose +
tubular +, acuminate +  and acute +
autoicous +  and sexual +
Patricia M. Eckel +
(M. Howe) W. R. Buck & D. H. Norris +
narrowing +
broadly decurrent;abruptly concave-auriculate +
Hedwigia albicans var. detonsa +
spreading +  and erect +
concave +, ovatelanceolate +, oblong-ovate +  and ovate +
stoloniform-flagelliform +
conic-mitrate +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.09 cm0.9 mm <br />9.0e-4 m <br />) +
redbrown +  and brown +
stegocarpous +
incurved +  and erect +
obovoid +, short-ovoid +, subglobose +  and turbinate-urceolate +
1 cm10 mm <br />0.01 m <br /> (?) +
smooth +  and simple-papillose +
short-rectangular +, quadrate +  and oblate +
enlarged +  and small +
differentiated +
low to moderate elevations (200-1500 m) +
River canyons, open forested lands, dry or moist acidic rock, sandstone, boulders and cliff faces near creeks, open, diffusely lighted mixed forests of Aesculus californica, Calocedrus, Pinus coulteri, Quercus chrysolepis, Torreya, and Umbellularia +
capillary +
imbricate +  and subsecund +
striolate +
broadly channeled;involute +
0.2 cm2 mm <br />0.002 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
ciliate +  and entire +
erect +  and incurved +
oblong-rhombic +, short-rectangular +  and subquadrate +
subumbonate-apiculate +
planoconvex +
multifid +  and simple +
few +  and many +
Capsules mature spring. +
truncate +  and sharp +
Nova Hedwigia +
strongly yellow +  and red-orange +
1 +  and several +
irregularly branched +  and subpinnate +
spreading +, erect-ascending +  and creeping +
flagelliform-tapered +  and stoloniform +
reddish-brown +
straight +
swelling;slender;stout +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.09 cm0.9 mm <br />9.0e-4 m <br />) +
Endemic +  and Illustrated +
flattened +  and spheroid +
secund +  and foliate +
spreading +  and erect +
concave +, ovatelanceolate +, oblong-ovate +  and ovate +
enlarged +  and small +
Hedwigia detonsa +
Hedwigia +
species +
yellow-hyaline +
long-filiform +
specialized +
paroicous +, autoicous +, sexual +  and asexual +
porose-sinuate +
enlarged +  and small +
pale buff +  and redbrown +