Heuchera caroliniana

(Rosendahl Butters & Lakela) E. F. Wells

Rhodora 81: 575. 1979,.

Common names: Carolina alum-root
Basionym: Heuchera americana var. caroliniana Rosendahl, Butters & Lakela Minnesota Stud. Pl. Sci. 2: 59. 1936
Treatment appears in FNA Volume 8. Treatment on page 95. Mentioned on page 84, 88.

Herbs acaulescent; caudex branched. Flowering-stems leafy, 30–105 cm, short-stipitate-glandular. Leaves: petiole glabrous or short-stipitate-glandular; blade broadly ovate, shallowly 5-lobed, 3–10.5 cm, base cordate or nearly truncate, lobes rounded or ovate, margins dentate, apex acute or obtuse, surfaces glabrous or minutely stipitate-glandular on veins abaxially, glabrous or short-stipitate-glandular adaxially. Inflorescences diffuse. Flowers: hypanthium weakly bilaterally symmetric, free 1.3–2.5 mm, green, subglobose, abruptly inflated distal to adnation to ovary, 2.8–4.5 mm, short-stipitate-glandular; sepals erect, green-tipped, equal, 0.4–1.5 mm, apex rounded; petals erect, white or pink, broadly spatulate, unlobed, 0.6–1 mm, margins entire; stamens exserted 0.2–1.5 mm; styles included from 1 mm to exserted 1.1 mm, 1–3.5 mm, to 0.1 mm diam; (stigmas from 0.5 mm included to 1.1 mm exserted). Capsules ovoid, 4–7 mm, beaks divergent, not papillose. Seeds dark-brown, ellipsoid, 0.5–0.8 mm. 2n = 14.


Phenology: Flowering Apr–Jun.
Habitat: Rich upland woods on base-saturated substrates, basic dikes and basic rock outcroppings
Elevation: 30-300 m

Discussion

The geographic range of Heuchera caroliniana is contiguous with and appears to be a southern extension of that of H. pubescens. Heuchera americana var. americana occurs in counties adjacent to the range of H. caroliniana but they are sympatric only along the boundary; intergradation between the two is infrequent. Mixed populations of H. caroliniana and H. americana var. americana have not been seen.

Selected References

None.

Lower Taxa

None.

"full" is not a number.

... more about "Heuchera caroliniana"
epigynous +, perigynous +  and hypogynous +
rounded +, obtuse +  and acute +
Elizabeth Fortson Wells +  and Barbara Greene Shipes +
(Rosendahl Butters & Lakela) E. F. Wells +
truncate +  and cordate +
Heuchera americana var. caroliniana +
not papillose +
dentate +, serrate +  and entire +
glandular-ciliate +  and ciliate +
5-lobed +  and ovate +
3 cm30 mm <br />0.03 m <br /> (10.5 cm105 mm <br />0.105 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
3-carpellate +  and 2-carpellate +
Carolina alum-root +
ebracteate +, bracteate +  and 2-300(-1000+)-flowered +
N.C. +, S.C. +  and Va. +
30-300 m +
unisexual +  and bisexual +
30 cm300 mm <br />0.3 m <br /> (105 cm1,050 mm <br />1.05 m <br />) +
2-3(-4)-beaked +  and folliclelike +
Rich upland woods on base-saturated substrates, basic dikes and basic rock outcroppings +
multicellular +
short-stipitate-glandular +
inflated +  and subglobose +
0.28 cm2.8 mm <br />0.0028 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
bracteate +, not secund +  and 100-1000-flowered +
dense +  and diffuse +
opposite +  and alternate +
ovate;rounded +
entire +  and dentate +
0.01 cm0.1 mm <br />1.0e-4 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
crassinucellate +, unitegmic +  and bitegmic +
epigynous +, perigynous +  and hypogynous +
pink +  and white +
distinct +
unlobed +  and spatulate +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
peltate +  and jointed +
short-stipitate-glandular +  and glabrous +
Flowering Apr–Jun. +
unequal +
dark-brown +
straight +
muricate +, cellular-rugulose +, tuberculate +, pitted +, papillate +  and wrinkled +
ellipsoid +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
green-tipped +
distinct +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
unbranched +  and branched +
persistent +
horizontal +  and erect +
0 cm0 mm <br />0 m <br /> (0.01 cm0.1 mm <br />1.0e-4 m <br />) +
connate +  and distinct +
exserted +  and included +
0.1 cm1 mm <br />0.001 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
short-stipitate-glandular +, glabrous +  and stipitate-glandular +
Heuchera caroliniana +
Heuchera +
species +
glandular-ciliate +  and ciliate +
crenate +  and dentate +
annual +, biennial +  and perennial +