Horkelia cuneata var. puberula

(Rydberg) Ertter & Reveal

Novon 17: 318. 2007.

Common names: Mesa horkelia
Conservation concernEndemic
Basionym: Horkelia puberula Rydberg Bull. Torrey Bot. Club 25: 55. 1898,
Synonyms: H. cuneata subsp. puberula (Rydberg) D. D. Keck P. kelloggii var. puberula (Rydberg) Hoover P. lindleyi var. puberula (Rydberg) Jepson
Treatment appears in FNA Volume 9. Treatment on page 255. Mentioned on page 254.

Plants green; eglandular hairs sparse or absent, spreading, glandular-hairs not obscured. Stems erect, 3–6 dm. Leaflets 5–12 per side, elliptic to obovate, 5–25 mm, distinctly pinnately veined. Inflorescences open, most flowers arranged individually. Pedicels (2–) 5–10 mm, proximalmost to 40 mm. Flowers: hypanthium interior rim usually glabrous, sometimes sparsely pilose; petals oblanceolate to narrowly obovate, 2–4 mm wide; filaments 1–2.5 × 0.5–2 mm.


Phenology: Flowering spring–summer.
Habitat: Coastal ranges away from the immediate coast, sandy to gravelly soil, coastal sage and chaparral communities
Elevation: 70–900 m

Discussion

Of conservation concern.

At its most distinctive, var. puberula is represented by historic collections from the hills and plains of Los Angeles, western Riverside, extreme southwestern San Bernardino, and northwestern San Diego counties; these populations have now largely been destroyed by urbanization. Scattered populations still exist within this core area, to what extent and with what level of ensured continuity remains uncertain. In addition, multiple populations of Horkelia cuneata from as far north and west as San Luis Obispo County have characteristics of var. puberula; these tend to intergrade with var. cuneata, as do some historic collections from the core area.

Selected References

None.

Lower Taxa

None.

"/3lengthofsepals" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property.

rugose +  and smooth +
1.5mm +  and 1.8mm +
longer than wide +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.11 cm1.1 mm <br />0.0011 m <br />) +
truncate +  and emarginate +
Barbara Ertter +  and James L. Reveal +
(Rydberg) Ertter & Reveal +
compound +  and simple +
opposite +  and alternate +
5 cm50 mm <br />0.05 m <br /> (10 cm100 mm <br />0.1 m <br />) +
4 cm40 mm <br />0.04 m <br /> (5 cm50 mm <br />0.05 m <br />) +
10 cm100 mm <br />0.1 m <br /> (30 cm300 mm <br />0.3 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (4 cm40 mm <br />0.04 m <br />) +
flattened +
Horkelia puberula +
30 cm300 mm <br />0.3 m <br /> (40 cm400 mm <br />0.4 m <br />) +
planar +  and cylindric +
2 cm20 mm <br />0.02 m <br /> (30 cm300 mm <br />0.3 m <br />) +
reduced +
free +  and distinct +
Mesa horkelia +
open to congested +
not +  and aggregated +
70–900 m +
lanceolate +  and ovate +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
flattened +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
1.2 cm12 mm <br />0.012 m <br /> (1.7 cm17 mm <br />0.017 m <br />) +
unisexual +  and bisexual +
not +  and aggregated +
obliquely ovoid +  and reniform +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Coastal ranges away from the immediate coast, sandy to gravelly soil, coastal sage and chaparral communities +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
glabrous +  and pilose +
0 +  and 1/2 +
0.1 cm1 mm <br />0.001 m <br /> (0.55 cm5.5 mm <br />0.0055 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.7 cm7 mm <br />0.007 m <br />) +
crassinucellate +
pilose +  and glabrous +
opposite +  and alternate +
marcescent +
winter-persistent +
separate +  and slightly overlapping +
2.5 cm25 mm <br />0.025 m <br /> (3 cm30 mm <br />0.03 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
sparsely pilose +  and glabrate or sericeous +
elliptic +  and obovate +
0.5 cm5 mm <br />0.005 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (?) +  and 1.5 cm15 mm <br />0.015 m <br /> (?) +
oblanceolate +
palmate +  and pinnate +
shallowly toothed +  and deeply divided +
inferior +  and superior +
clustered +, biseriate +  and superposed +
0.2 cm2 mm <br />0.002 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
recurved +  and reflexed +
0.5 cm5 mm <br />0.005 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0 (?) +  and 4 (?) +
free +  and distinct +
0.4 cm4 mm <br />0.004 m <br /> (0.8 cm8 mm <br />0.008 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
Flowering spring–summer. +
adnate +, free +, connate +  and distinct +
0 cm0 mm <br />0 m <br /> (4 cm40 mm <br />0.04 m <br />) +
pilose +  and glabrous +
not arillate +
persistent +
free +  and distinct +
less reflexed +  and spreading +
lanceolate +
0.4 cm4 mm <br />0.004 m <br /> (0.65 cm6.5 mm <br />0.0065 m <br />) +
Conservation concern +  and Endemic +
free +  and distinct +
branched +  and simple +
stramineous +, reddish +  and green +
1 +  and 10 +
palmate +  and pinnate +
deciduous +
distinct +
subterminal +
filiform +
rough-thickened +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
H. cuneata subsp. puberula +, P. kelloggii var. puberula +  and P. lindleyi var. puberula +
Horkelia cuneata var. puberula +
Horkelia cuneata +
variety +
10 +  and 15 +
oblanceolate +  and obovate +
inconspicuous +
enlarged +
hairy +  and inconspicuously conspicuously glandular +