Horkelia fusca var. capitata

(Lindley) M. Peck

Man. Pl. Oregon, 398. 1941.

Common names: Palouse horkelia
Endemic
Basionym: Horkelia capitata Lindley Edwards’s Bot. Reg. 23: sub plate 1997. 1837
Synonyms: H. fusca subsp. capitata (Lindley) D. D. Keck Potentilla capitata Greene P. douglasii var. capitata (Lindley) J. T. Howell
Treatment appears in FNA Volume 9. Treatment on page 261. Mentioned on page 259, 260, 262.

Stems 4–9 dm. Basal leaves green, 10–20 (–35) cm; leaflets 4–6 (–8) per side, cuneate to broadly obovate, 10–30 (–35) × 10–25 (–30) mm, 1/2 to as wide as long; divided 1/5–1/4 to midrib into 8–15 teeth, surfaces not obscured, ± sparsely short-hirsute to glabrate. Cauline leaves 3–6; leaflets of proximalmost 3–5 per side. Inflorescences dark reddish purple, densely congested, comprising no more than 1/6 of stem, composed of 5–30-flowered glomerules, glandular-hairs usually conspicuously red-septate; bracts acuminate-lobed, obscuring pedicels and flowers even at maturity. Flowers: epicalyx bractlets (2–) 3.5–5 (–6) mm; hypanthium 2–3 × 3–5 mm; petals 4–6.5 mm; filaments 0.5–1.5 mm, longer than wide, anthers 0.5–0.6 mm; styles 1–1.5 mm. Achenes 1.6–1.8 mm.


Phenology: Flowering summer.
Habitat: Dry meadow edges, in conifer woodlands, mainly on volcanic or granitic soil
Elevation: 100–2300 m

Discussion

As here circumscribed, var. capitata is found in meadow margins and grassy openings in the forested mountains surrounding the Palouse Prairie (in the broad sense) of northern Idaho, southeastern Washington, and northeastern Oregon. This is a significantly more limited range than accepted by D. D. Keck (1938) and in subsequent floras (for example, B. Ertter 1993d), with the bulk of what had been treated as var. (or subsp.) capitata here treated as var. pseudocapitata. The distinctions between the two varieties are more matters of degree than absolute differences, var. capitata generally having more congested, more purple, more glandular inflorescences in which the pedicels and hypanthia remain covered by larger involucral bracts divided into more elongate teeth. Plants of var. capitata can also be significantly larger, especially in the remnant prairie patches of northern Idaho, with more numerous cauline leaves having more pairs of larger leaflets. The extremes of the two varieties are reasonably distinct, with a significant biogeographic underpinning, but the differences merge in the Blue Mountains of Oregon.

Given the California focus of the published source (P. A. Munz 1959), the chromosome count of 2n = 28 for var. capitata is most likely based on material here treated as var. pseudocapitata.

Selected References

None.

Lower Taxa

None.

"/2lengthoftonearlyequaltosepals" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property.

1.6mm +  and 1.8mm +
longer than wide +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
pink +  and rose +
Barbara Ertter +  and James L. Reveal +
(Lindley) M. Peck +
compound +  and simple +
opposite +  and alternate +
20 cm200 mm <br />0.2 m <br /> (40 cm400 mm <br />0.4 m <br />) +
20 cm200 mm <br />0.2 m <br /> (35 cm350 mm <br />0.35 m <br />) +
4 cm40 mm <br />0.04 m <br /> (7 cm70 mm <br />0.07 m <br />) +
4 cm40 mm <br />0.04 m <br /> (20 cm200 mm <br />0.2 m <br />) +
10 cm100 mm <br />0.1 m <br /> (20 cm200 mm <br />0.2 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (4 cm40 mm <br />0.04 m <br />) +
flattened +
Horkelia capitata +
30 cm300 mm <br />0.3 m <br /> (40 cm400 mm <br />0.4 m <br />) +
planar +  and cylindric +
2 cm20 mm <br />0.02 m <br /> (30 cm300 mm <br />0.3 m <br />) +
acuminate-lobed +
reduced +
free +  and distinct +
Palouse horkelia +
open to congested +
Idaho +, Oreg. +  and Wash. +
not +  and aggregated +
100–2300 m +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.03 cm0.3 mm <br />3.0e-4 m <br />) +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
longer than wide +
flattened +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
unisexual +  and bisexual +
not +  and aggregated +
obliquely ovoid +  and reniform +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Dry meadow edges, in conifer woodlands, mainly on volcanic or granitic soil +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.55 cm5.5 mm <br />0.0055 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
crassinucellate +
opposite +  and alternate +
marcescent +
winter-persistent +
overlapping +  and separate +
3 cm30 mm <br />0.03 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
2.5 cm25 mm <br />0.025 m <br /> (3 cm30 mm <br />0.03 m <br />) +
1 cm10 mm <br />0.01 m <br /> (3 cm30 mm <br />0.03 m <br />) +
glabrate +, sparsely +  and more or less densely short-villous or hirsute +
divided +, cuneate +  and broadly obovate +
1 cm10 mm <br />0.01 m <br /> (?) +  and 2.5 cm25 mm <br />0.025 m <br /> (?) +
oblanceolate +
palmate +  and pinnate +
shallowly toothed +  and deeply divided +
inferior +  and superior +
clustered +, biseriate +  and superposed +
0.3 cm3 mm <br />0.003 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0 (?) +  and 4 (?) +
0.6 cm6 mm <br />0.006 m <br /> (0.65 cm6.5 mm <br />0.0065 m <br />) +
free +  and distinct +
0.4 cm4 mm <br />0.004 m <br /> (0.65 cm6.5 mm <br />0.0065 m <br />) +
Flowering summer. +
adnate +, free +, connate +  and distinct +
Man. Pl. Oregon, +
not arillate +
0.4 cm4 mm <br />0.004 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
persistent +
free +  and distinct +
spreading +  and more or less reflexed +
lanceolate +
0.2 cm2 mm <br />0.002 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
free +  and distinct +
branched +  and simple +
stramineous +, reddish +  and green +
ascending +  and erect +
/6 +, 1 +  and no +
palmate +  and pinnate +
deciduous +
distinct +
subterminal +
filiform +
rough-thickened +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
less sparsely short-hirsute +  and glabrate +
H. fusca subsp. capitata +, Potentilla capitata +  and P. douglasii var. capitata +
Horkelia fusca var. capitata +
Horkelia fusca +
variety +
5 +  and 15 +
oblanceolate +  and obovate +
inconspicuous +
enlarged +
grayish +, reddish +  and green +
hairy +  and inconspicuously conspicuously glandular +