Horkelia fusca var. pseudocapitata

(Rydberg ex Howell) M. Peck

Man. Pl. Oregon, 398. 1941.

Common names: Intermountain horkelia
Endemic
Basionym: Horkelia pseudocapitata Rydberg ex Howell Fl. N.W. Amer., 180. 1898
Synonyms: H. fusca subsp. pseudocapitata (Rydberg ex Howell) D. D. Keck Potentilla douglasii var. pseudocapitata (Rydberg ex Howell) Jepson
Treatment appears in FNA Volume 9. Treatment on page 262. Mentioned on page 259, 260, 261, 263.

Stems 3–6 dm. Basal leaves green, 10–35 (–40) cm; leaflets 4–6 (or 7) per side, narrowly to broadly obovate, 10–30 (–35) × (5–) 10–20 (–25) mm, 1/2 to nearly as wide as long, divided 1/5–1/4 to midrib into 8–14 teeth, surfaces not obscured, ± sparsely hirsute to glabrate. Cauline leaves 1–4 (or 5); leaflets of proximalmost 2 or 3 (–5) per side. Inflorescences green to reddish purple, congested to open, comprising 1/6–1/3 of stem, composed of 10–30-flowered glomerules, glandular-hairs not or obscurely red-septate; bracts acute-lobed, not obscuring pedicels and flowers at maturity. Flowers: epicalyx bractlets 2–3.5 (–4) mm; hypanthium 2 × 2.5–4 mm; petals (3–) 4–6 mm; filaments 0.5–1.5 mm, usually longer than wide, anthers 0.5–0.6 mm; styles 1–1.5 mm. Achenes 1.6–1.8 mm. 2n = 28.


Phenology: Flowering summer.
Habitat: Dry meadow edges, often with sagebrush, aspen, and/or willows, open conifer woodlands, mainly on volcanic or granitic soil
Elevation: 900–2300 m

Distribution

V9 416-distribution-map.jpg

Calif., Idaho, Nev., Oreg.

Discussion

The application of the epithet pseudocapitata here differs significantly from that of P. A. Rydberg (1908c), D. D. Keck (1938), and most floras, who used it for the taxon that is here called var. brownii. Instead, var. pseudocapitata encompasses the bulk of what had been called var. (or subsp.) capitata, except in mountains bordering the Palouse Prairie in Idaho, Oregon, and Washington. For nomenclatural details, see B. Ertter and J. L Reveal (2007).

As here circumscribed, var. pseudocapitata is a relatively large, big-petaled variety that grows in the mountains in and bordering the northern Intermountain Region in northeastern California (primarily the Warner Mountains), northern Nevada, southeastern Oregon, and southern Idaho, extending northward along valleys from Camas to Blaine counties. Intermediate plants are common where the range intersects those of var. brownii, var. capitata, and var. parviflora, though in the core of its range var. pseudocapitata is reasonably distinctive and uniform.

Selected References

None.

Lower Taxa

None.

"/2lengthoftonearlyequaltosepals" is not declared as a valid unit of measurement for this property."dm" is not declared as a valid unit of measurement for this property.

1.6mm +  and 1.8mm +
longer than wide +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
pink +  and rose +
Barbara Ertter +  and James L. Reveal +
(Rydberg ex Howell) M. Peck +
compound +  and simple +
opposite +  and alternate +
20 cm200 mm <br />0.2 m <br /> (40 cm400 mm <br />0.4 m <br />) +
35 cm350 mm <br />0.35 m <br /> (40 cm400 mm <br />0.4 m <br />) +
4 cm40 mm <br />0.04 m <br /> (7 cm70 mm <br />0.07 m <br />) +
4 cm40 mm <br />0.04 m <br /> (20 cm200 mm <br />0.2 m <br />) +
10 cm100 mm <br />0.1 m <br /> (35 cm350 mm <br />0.35 m <br />) +
1.5 cm15 mm <br />0.015 m <br /> (4 cm40 mm <br />0.04 m <br />) +
flattened +
Horkelia pseudocapitata +
30 cm300 mm <br />0.3 m <br /> (40 cm400 mm <br />0.4 m <br />) +
planar +  and cylindric +
2 cm20 mm <br />0.02 m <br /> (30 cm300 mm <br />0.3 m <br />) +
acute-lobed +
reduced +
free +  and distinct +
Intermountain horkelia +
open to congested +
Calif. +, Idaho +, Nev. +  and Oreg. +
not +  and aggregated +
900–2300 m +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.03 cm0.3 mm <br />3.0e-4 m <br />) +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
usually longer than wide +
flattened +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (1.2 cm12 mm <br />0.012 m <br />) +
unisexual +  and bisexual +
not +  and aggregated +
obliquely ovoid +  and reniform +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
Dry meadow edges, often with sagebrush, aspen, and/or willows, open conifer woodlands, mainly on volcanic or granitic soil +
2 cm20 mm <br />0.02 m <br /> (?) +
0.1 cm1 mm <br />0.001 m <br /> (0.55 cm5.5 mm <br />0.0055 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
congested +  and open +
green +  and reddish purple +
crassinucellate +
opposite +  and alternate +
marcescent +
winter-persistent +
overlapping +  and separate +
3 cm30 mm <br />0.03 m <br /> (3.5 cm35 mm <br />0.035 m <br />) +
2 cm20 mm <br />0.02 m <br /> (2.5 cm25 mm <br />0.025 m <br />) +
1 cm10 mm <br />0.01 m <br /> (3 cm30 mm <br />0.03 m <br />) +
glabrate +, sparsely +  and more or less densely short-villous or hirsute +
divided +  and obovate +
1 cm10 mm <br />0.01 m <br /> (?) +  and 2 cm20 mm <br />0.02 m <br /> (?) +
oblanceolate +
palmate +  and pinnate +
shallowly toothed +  and deeply divided +
inferior +  and superior +
clustered +, biseriate +  and superposed +
0.3 cm3 mm <br />0.003 m <br /> (1 cm10 mm <br />0.01 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0 (?) +  and 4 (?) +
0.3 cm3 mm <br />0.003 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
free +  and distinct +
0.4 cm4 mm <br />0.004 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
Flowering summer. +
adnate +, free +, connate +  and distinct +
Man. Pl. Oregon, +
not arillate +
0.4 cm4 mm <br />0.004 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
persistent +
free +  and distinct +
spreading +  and more or less reflexed +
lanceolate +
0.2 cm2 mm <br />0.002 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
free +  and distinct +
branched +  and simple +
stramineous +, reddish +  and green +
ascending +  and erect +
1/6 +  and 1/3 +
palmate +  and pinnate +
deciduous +
distinct +
subterminal +
filiform +
rough-thickened +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
not obscured +
less sparsely hirsute +  and glabrate +
H. fusca subsp. pseudocapitata +  and Potentilla douglasii var. pseudocapitata +
Horkelia fusca var. pseudocapitata +
Horkelia fusca +
variety +
8 +  and 14 +
oblanceolate +  and obovate +
inconspicuous +
enlarged +
grayish +, reddish +  and green +
hairy +  and inconspicuously conspicuously glandular +