Ivesia gordonii var. alpicola

(Rydberg ex Howell) Ertter & Reveal

Novon 17: 322. 2007.

Common names: Howell’s ivesia
Endemic
Basionym: Ivesia alpicola Rydberg ex Howell Fl. N.W. Amer., 182. 1898
Synonyms: Horkelia gordonii var. alpicola (Rydberg ex Howell) Rydberg
Treatment appears in FNA Volume 9. Treatment on page 234. Mentioned on page 235.

Stems usually greenish, rarely reddish, usually ascending to erect, sometimes decumbent, 0.3–2 dm, not or sparsely hirsute to villous, glandular-puberulent or pubescent. Basal leaves 2–8 (–10) × 0.5–1 (–1.5) cm; leaflets 2–5 (–9) mm, hirsute to pilose, rarely minutely glandular at least marginally. Inflorescences 5–25-flowered, usually simple, 1–3 (–4) cm diam., glomerules usually 1, ± capitate. Flowers 5–9 mm diam.; epicalyx bractlets (1.5–) 2–3 (–4) mm; hypanthium (2–) 2.5–3.5 (–4) × 2.5–4 mm; sepals 2–4 (–5) mm, acute; anthers rarely red-margined.


Phenology: Flowering summer.
Habitat: Dry grassy areas on talus, or gravelly or rocky slopes and ridges, in subalpine and alpine montane conifer woodlands, alpine tundra
Elevation: 2100–3300 m

Distribution

V9 359-distribution-map.jpg

Calif., Idaho, Mont., Oreg., Wash.

Discussion

P. A. Rydberg (1898) used var. alpicola (as Horkelia gordonii var. alpicola) for compact, high-elevation plants throughout the range of Ivesia gordonii. As circumscribed here, var. alpicola is restricted to plants occurring disjunctly in an arc around the northwestern part of the species range. Populations occur in the northern Coast Ranges and central Sierra Nevada of California, sporadically to Mount Adams and the Wenatchee Mountains in Washington, and from there to the mountains of central Idaho and the Bitterroot Range of Montana. The variety is also disjunct on the Owyhee Mountains of southwestern Idaho.

Plants of var. alpicola average shorter than those of var. gordonii, which occurs farther east and south. In var. alpicola, stems with only glandular indumentum are common; such stems are rare in var. gordonii. Unlike those of var. gordonii, the leaflets of var. alpicola are frequently glandular with only marginal hairs.

Selected References

None.

Lower Taxa

None.

"dm" is not declared as a valid unit of measurement for this property.

not carunculate +
grayish brown +  and mottled brown +
longer than wide +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
Barbara Ertter +  and James L. Reveal +
(Rydberg ex Howell) Ertter & Reveal +
compound +  and simple +
opposite +  and alternate +
8 cm80 mm <br />0.08 m <br /> (10 cm100 mm <br />0.1 m <br />) +
20 cm200 mm <br />0.2 m <br /> (25 cm250 mm <br />0.25 m <br />) +
1 cm10 mm <br />0.01 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
2 cm20 mm <br />0.02 m <br /> (8 cm80 mm <br />0.08 m <br />) +
3 cm30 mm <br />0.03 m <br /> (20 cm200 mm <br />0.2 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (1 cm10 mm <br />0.01 m <br />) +
Ivesia alpicola +
20 cm200 mm <br />0.2 m <br /> (25 cm250 mm <br />0.25 m <br />) +
narrowly oblong +  and filiform planar +
reduced +
0.5 cm5 mm <br />0.005 m <br /> (20 cm200 mm <br />0.2 m <br />) +
reduced +
elongate +
free +  and distinct +
branched +  and simple +
Howell’s ivesia +
Calif. +, Idaho +, Mont. +, Oreg. +  and Wash. +
not +  and aggregated +
2100–3300 m +
0.3 cm3 mm <br />0.003 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0.13 cm1.3 mm <br />0.0013 m <br /> (0.25 cm2.5 mm <br />0.0025 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
unisexual +  and bisexual +
not +  and aggregated +
deciduous +
obliquely ovoid +  and reniform +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
loose +  and capitate +
Dry grassy areas on talus, or gravelly or rocky slopes and ridges, in subalpine and alpine montane conifer woodlands, alpine tundra +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
0.3 cm3 mm <br />0.003 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
0.4 cm4 mm <br />0.004 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.35 cm3.5 mm <br />0.0035 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
simple +  and 5-25-flowered +
1 cm10 mm <br />0.01 m <br /> (3 cm30 mm <br />0.03 m <br />) +
crassinucellate +
opposite +  and alternate +
winter-marcescent +
deciduous +  and persistent +
0.5 cm5 mm <br />0.005 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
, +  and distinct +
not +  and distinguishable +
hirsute +  and pilose +
0.2 cm2 mm <br />0.002 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
narrowly oblanceolate +  and obovate +
verticillate +
toothed +, unlobed +  and lobed +
inferior +  and superior +
clustered +, biseriate +  and superposed +
0.3 cm3 mm <br />0.003 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0 (?) +  and 4 (?) +
0.1 cm1 mm <br />0.001 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
free +  and distinct +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
0.5 cm5 mm <br />0.005 m <br /> (8 cm80 mm <br />0.08 m <br />) +
Flowering summer. +
adnate +, free +, connate +  and distinct +
not arillate +
0.4 cm4 mm <br />0.004 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
persistent +
free +  and distinct +
erect +  and spreading +
0.2 cm2 mm <br />0.002 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
free +  and distinct +
branched +  and simple +
reddish +  and greenish +
usually ascending +  and erect +
2 +  and 20 +
palmate +  and pinnate +
0.45 cm4.5 mm <br />0.0045 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
deciduous +
distinct +
subterminal +
filiform +
rough-thickened +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.45 cm4.5 mm <br />0.0045 m <br />) +
Horkelia gordonii var. alpicola +
not fleshy +
Ivesia gordonii var. alpicola +
Ivesia gordonii +
variety +
inconspicuous +
stipelike +, elongate +, less flat +  and conic or turbinate +
enlarged +
glabrate +  and sparsely densely glandular-puberulent +