Ivesia sect. Comarella

(Rydberg) Ertter & Reveal

Novon 17: 316. 2007.

Conservation concernEndemic
Basionym: Undefined sect. Comarella Rydberg Monogr. N. Amer. Potentilleae, 156, plates 96, 97, figs. 1–5. 1898
Treatment appears in FNA Volume 9. Treatment on page 243. Mentioned on page 219, 221, 271.

Plants usually ± tufted, not forming hanging clumps, not aromatic; taproot stout, not fusiform or fleshy. Stems (1.8–) 2–6 (–6.5) dm. Basal leaves cylindric; stipules usually present, sometimes much-reduced; leaflets loosely to ± tightly overlapping, individually distinguishable at least distally, toothed 1/3–1/2 or lobed to base, sometimes entire, sparsely to ± densely hairy, sometimes glabrate; terminal leaflets indistinct. Cauline leaves 1–3, not paired; blade well developed. Inflorescences open, flowers arranged individually. Pedicels remaining straight. Flowers: hypanthium patelliform; petals medially reflexed, yellow or red, not clawed, apex acute to obtuse; stamens 5, anthers longer than wide, laterally dehiscent; carpels 1–5. Achenes horizontal, smooth, not carunculate.

Distribution

sw United States

Discussion

Species 2 (2 in the flora).

Rydberg established sect. Comarella to accommodate two species with wheel-shaped centrally turbinate hypanthia, linear petals, five stamens, and few carpels that are nearly horizontal at maturity. He understood both species to have purple petals, but the petals of Ivesia sabulosa are actually bright yellow. Other than flower color and leaflet lobing, the two species are nearly identical, with diffuse inflorescences on erect stems emerging from tufts of erect, slender leaves with numerous pairs of leaflets. The medially reflexed petals are unique to these two species of Ivesia, though not always well developed. The stipules of the basal leaves are often little more than slightly rounded, winglike extensions of the petiole base, if they are present at all.

Comarella is the only section of the genus without representation in California. The plants occur at the southeastern corner of the range of Ivesia in the southern Great Basin, southwestern Colorado Plateau, and adjacent Mojave Desert. The Grand Canyon of the Colorado River divides the range of the two species in northern Arizona.

Selected References

None.

Key

1 Petals brick- to blood-red; leaflets incised 1/3–1/2 to base; s of Colorado River, Arizona. Ivesia multifoliolata
1 Petals yellow; leaflets incised to base; n of Colorado River, Arizona, Nevada, Utah. Ivesia sabulosa

"dm" is not declared as a valid unit of measurement for this property.

... more about "Ivesia sect. Comarella"
not carunculate +
longer than wide +
Barbara Ertter +  and James L. Reveal +
(Rydberg) Ertter & Reveal +
compound +  and simple +
opposite +  and alternate +
Undefined sect. Comarella +
20 cm200 mm <br />0.2 m <br /> (25 cm250 mm <br />0.25 m <br />) +
narrowly oblong +  and filiform planar +
0.5 cm5 mm <br />0.005 m <br /> (20 cm200 mm <br />0.2 m <br />) +
reduced +
elongate +
free +  and distinct +
branched +  and simple +
sw United States +
not +  and aggregated +
0.4 cm4 mm <br />0.004 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
unisexual +  and bisexual +
not +  and aggregated +
deciduous +
obliquely ovoid +  and reniform +
0.08 cm0.8 mm <br />8.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
loose +  and capitate +
0.3 cm3 mm <br />0.003 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
crassinucellate +
opposite +  and alternate +
winter-marcescent +
deciduous +  and persistent +
, +  and distinct +
distinguishable +
glabrate +  and hairy +
lobed +  and toothed +
oblanceolate +  and oval or obovate +
verticillate +
toothed +, unlobed +  and lobed +
inferior +  and superior +
clustered +, biseriate +  and superposed +
0 (?) +  and 4 (?) +
Red (?) +  and Yellow (?) +
free +  and distinct +
adnate +, free +, connate +  and distinct +
not arillate +
persistent +
free +  and distinct +
erect +  and spreading +
lanceolate +  and broadly ovate or deltate +
Conservation concern +  and Endemic +
free +  and distinct +
branched +  and simple +
reddish +, grayish +  and green +
pendent +  and erect +
2 +  and 20 +
palmate +  and pinnate +
much-reduced +
deciduous +
distinct +
subterminal +
filiform +
rough-thickened +
Horkelia subg. Ivesia +, Potentilla sect. Ivesia +  and Potentilla subg. Ivesia +
not fusiform +
Ivesia sect. Comarella +
section +
inconspicuous +
stipelike +, elongate +, less flat +  and conic or turbinate +
enlarged +
glabrate +  and sparsely densely glandular-puberulent +