Ludwigia suffruticosa

Walter

Fl. Carol., 90. 1788.

Common names: Shrubby primrose-willow or seedbox
Synonyms: Isnardia suffruticosa (Walter) Kuntze
Treatment appears in FNA Volume 10.

Herbs with 1–3 rhizomes 0.6–5.5 × 0.2–0.5 mm, often branched, glabrous or densely hirtellous, sometimes also form­ing stolons 8–80 cm, 1.1–2.2 mm thick, branched, glabrous or sparsely to densely hirtellous. Stems erect, slightly ridged, unbranched or slightly branched, (16–) 30–90 cm, glabrous or strigillose to hirtellous, especially on distal parts. Leaves alternate; stipules deltate, 0.25–0.45 × 0.15–0.4 mm; rhizomes: sessile, blades minute, appressed, and scalelike, oblate or suborbiculate, 0.3–0.6 × 0.6–0.9 mm; stolons: petiole 0.1–0.6 cm, blade oblong or oblanceolate-elliptic to spatulate, 0.4–3.5 × 0.2–1.5 cm; main-stem: sessile, blade lanceolate-elliptic or lanceolate-linear to linear, 2.5–9.5 × (0.1–) 0.3–0.9 cm, proximal ones shorter and often oblong or oblong-lanceolate, base rounded or obtuse, margins entire with obscure hydathodal glands, apex acuminate to acute, surfaces glabrous or, sometimes, pilose on proximal blades; bracts very reduced. Inflorescences densely clustered, terminal racemes or spikes, 1–5 (–12) cm; bracteoles attached at base of ovary or on pedicel distally, narrowly lanceolate, 3.5–5 (–6) × (1.2–) 1.4–2 mm, surfaces usually glabrous, sometimes pilose abaxially. Flowers: sepals ascending, pale green or white adaxially, broadly ovate-deltate, 2.3–3.5 (–4) × 2.3–3.2 (–3.8) mm, margins entire, apex acuminate, surfaces glabrous; petals 0; filaments yellow or cream, 1.2–2 mm, distinctly dilated toward base, anthers 0.7–1 (–1.3) × 0.5–0.7 mm; pollen shed in tetrads; ovary broadly obovoid or cupshaped, 2.2–3 × 2.3–3.3 mm; nectary disc elevated 0.5–0.6 mm on ovary apex, pale-yellow, 1.8–3.1 mm diam., obscurely 4-lobed, glabrous; style pale-yellow, 0.9–1.7 mm, gla­brous, stigma pale green to white, globose to capi­tate, 0.4–0.8 × 0.5–0.8 mm, distinctly 4-lobed, not exserted beyond anthers. Capsules broadly obpyramidal, angles rounded, sometimes subspherical, 2.5–4.3 ×2.5–4.5 (–5) mm, hard-walled, dehiscent by apical ring, pedicel 0.5–1.5 (–2) mm. Seeds brown, elliptic-oblong, curved on both ends, 0.5–0.6 × 0.2–0.3 mm, surface cells ± isodiametric. 2n = 32.


Phenology: Flowering May–Sep.
Habitat: Sandy ditches, marshes, wet meadows, limestone sinks, cypress swamps, moist pinelands.
Elevation: 0–150 m.

Distribution

Ala., Fla., Ga., N.C., S.C.

Discussion

Ludwigia suffruticosa is distinctive by virtue of its highly condensed inflorescence and sessile leaves. It also differs from other species in sect. Isnardia in that it perennates mainly by underground rhizomes. This apetalous species has showy bracts and attracts many insects, including bumblebees, wasps, and honeybees (C. I. Peng 1989). Its center of distribution is in Florida, extending along the coastal plain barely to Alabama on the west and barely to southern North Carolina on the northeast. Recent reports of this species from Mississ­ippi and from Mexico (Chiapas and Oaxaca) have not been confirmed.

Selected References

None.

Lower Taxa

None.
... more about "Ludwigia suffruticosa"
0.45 cm4.5 mm <br />0.0045 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
0.25 cm2.5 mm <br />0.0025 m <br /> (0.43 cm4.3 mm <br />0.0043 m <br />) +
subspherical +  and rounded +
2.5mm +  and 4.5mm +
0.1 cm1 mm <br />0.001 m <br /> (0.13 cm1.3 mm <br />0.0013 m <br />) +
3(-5)-aperturate +
0.07 cm0.7 mm <br />7.0e-4 m <br /> (0.1 cm1 mm <br />0.001 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.07 cm0.7 mm <br />7.0e-4 m <br />) +
acuminate +  and acute +
Peter C. Hoch +
Walter +
obtuse +  and rounded +
Isnardia +
indehiscent +  and loculicidal +
0.1 cm1 mm <br />0.001 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
2.5 cm25 mm <br />0.025 m <br /> (9.5 cm95 mm <br />0.095 m <br />) +
lobed +, toothed +  and entire +
lanceolate-linear +  and linear +
0.3 cm3 mm <br />0.003 m <br /> (0.9 cm9 mm <br />0.009 m <br />) +
reduced +
0.5 cm5 mm <br />0.005 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.14 cm1.4 mm <br />0.0014 m <br />) +
dark red +  and black +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.5 cm5 mm <br />0.005 m <br />) +
lanceolate +
0.14 cm1.4 mm <br />0.0014 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
straight +  and slightly curved +
spreading +  and erect +
obpyramidal +
Shrubby primrose-willow or seedbox +
Ala. +, Fla. +, Ga. +, N.C. +  and S.C. +
0–150 m. +
cream +  and yellow +
basifixed +  and versatile +
0.12 cm1.2 mm <br />0.0012 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
long-pedicellate +  and subsessile +
straight +  and slightly curved +
by a terminal pore +  and dehiscent +
spreading +  and erect +
cylindric +  and clavate turbinate obpyramidal or globose +
Sandy ditches, marshes, wet meadows, limestone sinks, cypress swamps, moist pinelands. +
perennial +
hirtellous +  and glabrous +
deciduous +
0.18 cm1.8 mm <br />0.0018 m <br /> (0.31 cm3.1 mm <br />0.0031 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
0.22 cm2.2 mm <br />0.0022 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
cup-shaped +  and obovoid +
2.3mm +  and 3.3mm +
numerous +  and 1 +
submerged +
swollen +
0.15 cm1.5 mm <br />0.0015 m <br /> (0.2 cm2 mm <br />0.002 m <br />) +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.15 cm1.5 mm <br />0.0015 m <br />) +
5 +  and 4 +
0.1 cm1 mm <br />0.001 m <br /> (0.6 cm6 mm <br />0.006 m <br />) +
Flowering May–Sep. +
oblong-lanceolate +  and oblong +
Fl. Carol., +
paired +, solitary +  and cluster +
ascending +, decumbent +  and erect +
inconspicuous +
expanded +
0.06 cm0.6 mm <br />6.0e-4 m <br /> (0.55 cm5.5 mm <br />0.0055 m <br />) +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.05 cm0.5 mm <br />5.0e-4 m <br />) +
basal +  and cauline +
clavate +  and globose +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.06 cm0.6 mm <br />6.0e-4 m <br />) +
50 +  and 400 +
pitted +  and smooth +
elliptic-oblong +
0.02 cm0.2 mm <br />2.0e-4 m <br /> (0.03 cm0.3 mm <br />3.0e-4 m <br />) +
0.35 cm3.5 mm <br />0.0035 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
0.32 cm3.2 mm <br />0.0032 m <br /> (0.38 cm3.8 mm <br />0.0038 m <br />) +
white +  and pale green +
persistent +
post-anthesis +
5 +  and 4 +
ovate-deltate +
0.23 cm2.3 mm <br />0.0023 m <br /> (0.32 cm3.2 mm <br />0.0032 m <br />) +
parenchymatous +
parietal +, axile +  and placentation +
2 +  and 1 +
unequal +  and subequal +
2 times as many as sepals +
aquatic +, amphibious +  and terrestrial +
lenticular +
2 times as many as sepals +
branched +  and unbranched +
16 cm160 mm <br />0.16 m <br /> (30 cm300 mm <br />0.3 m <br />) +
sprawling +, decumbent +  and prostrate +
strigillose +  and hirtellous +
30 cm300 mm <br />0.3 m <br /> (90 cm900 mm <br />0.9 m <br />) +
pale green +  and white +
0.04 cm0.4 mm <br />4.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
4-lobed +  and globose +
0.05 cm0.5 mm <br />5.0e-4 m <br /> (0.08 cm0.8 mm <br />8.0e-4 m <br />) +
sessile +  and petiolate +
green +  and dark reddish +
0.025 cm0.25 mm <br />2.5e-4 m <br /> (0.045 cm0.45 mm <br />4.5e-4 m <br />) +
intrapetiolar +
0.015 cm0.15 mm <br />1.5e-4 m <br /> (0.04 cm0.4 mm <br />4.0e-4 m <br />) +
glabrous or +  and sparsely densely hirtellous +
8 cm80 mm <br />0.08 m <br /> (80 cm800 mm <br />0.8 m <br />) +
0.11 cm1.1 mm <br />0.0011 m <br /> (0.22 cm2.2 mm <br />0.0022 m <br />) +
pale-yellow +
0.09 cm0.9 mm <br />9.0e-4 m <br /> (0.17 cm1.7 mm <br />0.0017 m <br />) +
aquatic +, amphibious +  and terrestrial +
glabrous +, pilose +  and , +
papillate +
Isnardia suffruticosa +
Ludwigia suffruticosa +
Ludwigia sect. Isnardia +
species +
5 cm50 mm <br />0.05 m <br /> (12 cm120 mm <br />0.12 m <br />) +
1 cm10 mm <br />0.01 m <br /> (5 cm50 mm <br />0.05 m <br />) +
5 cm50 mm <br />0.05 m <br /> (12 cm120 mm <br />0.12 m <br />) +
1 cm10 mm <br />0.01 m <br /> (5 cm50 mm <br />0.05 m <br />) +
3(-5)-aperturate +