Malacomeles

(Decaisne) Decaisne

Ann. Gén. Hort. 23: 156. 1882.

Etymology: Greek malakos, soft, and melon, apple
Basionym: Cotoneaster sect. Malacomeles Decaisne Nouv. Arch. Mus. Hist. Nat. 10: 177. 1874,
Treatment appears in FNA Volume 9. Treatment on page 645. Mentioned on page 428, 646, 662.

Shrubs, 10–30 [–50] dm. Stems 1–20, ascending, much branched, often ± twisted, young stems terete, closely villous, smooth; bark brown or gray, young stems not hairy; long-shoots present, short-shoots sometimes present; unarmed; grayish or whitish-tomentose, sometimes glabrate; overwintering buds tomentose. Leaves drought-deciduous or persistent, cauline, simple, slightly conduplicate; stipules tardily deciduous or ± persistent, free, lanceolate, glandlike, margins entire; petiole present; blade elliptic or oblong-ovate to orbiculate, 0.5–1.5 [–3.1] cm, leathery, margins flat, denticulate or entire, teeth gland-tipped, venation pinnate, abaxial surface tomentose, adaxial ± glabrous. Inflorescences terminal on leafy shoots of season, 1–5 [–7] -flowered, corymbs [panicles], axes glabrous or hairy; bracts absent; bracteoles sometimes present. Pedicels present. Flowers: perianth and androecium epigynous, 10 mm diam.; hypanthium campanulate [urceolate], (2–) 3–4 mm, distally copiously white-tomentose; sepals 5, spreading, nearly orbiculate (inner broadly deltate); petals 5, white, round or kidney-shaped, base short-clawed; stamens [8–] 14–20 [–22], longer or shorter than petals; carpels 2 or 3 [–5], barely connate or distinct, adnate to proximal 1/2–4/5 of hypanthium, ovary appearing 4–10-loculed by false partitions, densely hairy, styles 2 or 3, ventral, distinct; ovules 2 (separated by false partition). Fruits berrylike pomes, translucent, vivid pink, drying purplish black, globose, 6–10 mm, glabrous; flesh whitish; hypanthium persistent; sepals persistent, erect or recurving; carpels cartilaginous; styles persistent. Seeds 4–6 (–10).

Distribution

United States, Mexico, Central America

Discussion

Species 2 (1 in the flora).

Malacomeles shares with Amelanchier and Peraphyllum the feature of false partitions within the carpels; the partitions partially divide the locules and make the fruit appear to have twice as many locules as the number of carpels (G. N. Jones 1945; K. R. Robertson et al. 1991; J. R. Rohrer et al. 1991). Sequences from multiple chloroplast and nuclear genes indicate that Malacomeles is closely related to Amelanchier and Peraphyllum (C. S. Campbell et al. 2007).

The other species in the genus, Malacomeles nervosa (Decaisne) G. N. Jones, is known from Nuevo León, Mexico south to Guatemala.

Selected References

None.

... more about "Malacomeles"
10 cm100 mm <br />0.1 m <br /> (?) +
Christopher S. Campbell +
(Decaisne) Decaisne +
hairy +  and glabrous +
gray +  and brown +
compound +  and simple +
opposite +  and alternate +
short-clawed +
Cotoneaster sect. Malacomeles +
1.5 cm15 mm <br />0.015 m <br /> (3.1 cm31 mm <br />0.031 m <br />) +
oblong-ovate +  and orbiculate +
0.5 cm5 mm <br />0.005 m <br /> (1.5 cm15 mm <br />0.015 m <br />) +
overwintering +
tomentose +
adnate +, distinct +  and connate +
United States +, Mexico +  and Central America +
not +  and aggregated +
Greek malakos, soft, and melon, apple +
unisexual +  and bisexual +
purplish black +  and vivid pink +
globose +
0.6 cm6 mm <br />0.006 m <br /> (1 cm10 mm <br />0.01 m <br />) +
not +  and aggregated +
not +  and aggregated +
0.2 cm2 mm <br />0.002 m <br /> (0.3 cm3 mm <br />0.003 m <br />) +
white-tomentose +
0.3 cm3 mm <br />0.003 m <br /> (0.4 cm4 mm <br />0.004 m <br />) +
crassinucellate +
persistent +  and drought-deciduous +
denticulate +
4-10-loculed +
inferior +  and superior +
biseriate +  and clustered +
collateral +  and basal +
10 cm100 mm <br />0.1 m <br /> (?) +
0 (?) +  and 4 (?) +
free +  and distinct +
adnate +, free +, connate +  and distinct +
Ann. Gén. Hort. +
not arillate +
persistent +
free +  and distinct +
recurving +, erect +  and spreading +
orbiculate +
glabrate +  and whitish-tomentose +
free +  and distinct +
twisted +  and branched +
1 +  and 20 +
persistent +  and deciduous +
persistent +
distinct +
elongate +
Malacomeles +
Rosaceae tribe Gillenieae +
gland-tipped +
inconspicuous +
17 +, 15 +, 9 +  and 8 +